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Term Definition
Caducous Falling off very early.
Abaxial The lower part of a structure such as a leaf, i.e., that part of a structure that faces away the main axis to which it is attached. For example, if a leaf is pushed upward and against a stem the surface facing away from the stem is the abaxial surface. Floral development of Bertholletia excelsa
Adaxial The upper part of a structure such as a leaf, i.e., that part of the structure that faces toward the main axis to which it is attached. For example, if a leaf is pushed upward and against a stem the surface facing toward the stem is the abaxial surface. Floral development of Bertholletia excelsa
Admedial Toward the axis of any structure, especially used to describe leaf venation. Venation of Eschweilera baguensis. Photo by S. A. Mori.
Alate Refers to a band of tissue running the length of a structure; = winged. Alate immature fruit of Gustavia brachycarpa. Photo by S. A. Mori
Androecial appendages In the Lecythidaceae, any prolongation from the surface of the androecium; if there are fertile anthers on the appendage they are called a stamens, if there are normal appearing anthers that produce only fodder pollen they are called staminodes, if there are only rudimentary anthers or no anthers at all the appendages they are called vestigial stamens, and if they produce nectar they are called nectar-producing vestigial stamens. Androecial hood: Lecythis pisonis-type. Photos by S. A. Mori
Androecial hood In zygomorphic-flowered Lecythidaceae with ligules, the distal part of the ligule that has become modified either by the outgrowth of appendices (= staminodes, vestigial stamens, or nectar-producing vestigial stamens), by an abrupt change in direction of the ligule, or by both. Androecial hoods are usuallly separated from the staminal ring by an appendage-free part of the ligule (the only known exception is and perhaps a few species of the Lecythis sec. Pisonis group. Androecial hoods of Lecythidaceae vary according to their degree of structural complexity and the type of androecial appendages they possess. The androecium of a zygomorphic-flowered Lecythidaceae. Photo by C. A. Gracie.
Androecial hood appendages swept inward A androecial hood type in which appendages arise from an expanded apical or subapical part of the ligule from where they curve into the flower but do not form a coil (e.g., as in species of ). The appendices are always more or less fused together at their bases. If the expanded part of the hood is subapical and the ligule extends beyond the thickening, the extension is called the anterior ligular extension. Do not confuse this term with curved appendages (see definition) Androecial hood: Bertholletia-type. Drawing by Bobbi Angell and photo by S. A. Mori
Androecial hood double-coiled The androecial hood is coiled twice inward. Medial section of Eschweilera atropetiolata flowers. Photo by C. Gracie.
Androecial hood flat The androecial hood is not curved or coiled inward at all Androecial hood: Lecythis pisonis-type. Photos by S. A. Mori
Androecial hood flat, thick The androecial hood, as seen in medial longitudinal section, is thickened dorsiventrally. Androecial hood: Lecythis corrugata-type. Drawing by B. angell.
Androecial hood flat, thin The androecial hood is not curved or coiled inward and it is not thickened as viewed in a medial longitudinal section. Androecial hood: Lecythis pisonis-type. Drawing by Bobbi Angell.
Androecial hood horizontally oriented (= hood horizontally oriented) An androecial hood with the first coil oriented at more-or-less 90 degrees to the main axis of the flower. Horizontally androecial hood orientation of Eschweilera apiculata. Photo by S. A. Mori.
Androecial hood obliquely oriented (= hood obliquely orientated) An androecial hood that is oriented at an angle less than 90 degrees to the main axis of the flower. Androeial hood orientation of Eschweilera calyculata. Photo by M. Stapf.
Androecial hood once-coiled The androecial hood turns inward once Medial section of the androecium of Lecythis tuyrana. Photo by C. Galdames (SCZ)
Androecial hood split A separation of the androecial hood into a posterior hood extension and an anterior hood extension. Medial sections sections of an androecium of Bertholletia excelsa. Photo by N. P. Smith.
Androecial hood triple-coiled The androecial hood makes three turns inward Androecium of Eschweilera pedicellata. Photo by S. A. Mori.
Androecial hood: Bertholletia-type The Bertholletia-type of androecial hood has appendages that are swept back into the interior of the hood which are called vestigial stamens; these appendages do not have anthers. Androecial hood: Bertholletia-type. Drawing by Bobbi Angell and photo by S. A. Mori
Androecial hood: Corythophora alta-type A thickened hood, bearing staminodes. Androecial hood: Corythophora alta-type. Photos by S. A. Mori.
Androecial hood: Couratari-type The androecial hood has an extra external flap found only in species of . Androecial hood o Couratari stellata. Photo by C. Gracie and drawing by B. Angell.
Androecial hood: Couroupita guianensis-type In this type of hood, there is an appendage-free area between the staminal ring and the androecial hood proper. The appendages bear normal looking anthers but they contain fodder pollen (=staminodes) and the hood is slightly thickened. Androecial hood of Couroupita guianensis. Drawing by Bobbi Angell.
Androecial hood: Couroupita subsessilis-type This type of androecial hood has three features: 1) an invagination, 2) a notch on the distal end as viewed from above, and 3) staminodes. Androecial hood: Couroupita subsessilis-type. Drawing by B. Angell.
Androecial hood: Eschweilera-type In this type of hood, the interior of the hood forms an inward coil and the vestigial stamens at the end of the coil are nectar-producing. Androecial hood: Eschweilera-type. Drawing by B. Angell.
Androecial hood: Lecythis corrugata-type In this type of androecial hood, the hood is flat and thickened, there are no appendages with anthers on the hood, and the vestigial stamens are poorly developed. Androecial hood: Lecythis corrugata-type. Drawing by B. angell.
Androecial hood: Lecythis idatimon-type The androecial hood is flat and there are only vestigial stamens representing the appendages of the hood. Androecial hood: Lecythis idatimon-type. Drawing by B. Angell. Photo by C. Gracie.
Androecial hood: Lecythis ollaria-type This type of androecial hood turns inward but does not form a complete coil, the vestigial stamens are all the same size, and lack anthers; thus, they are called vestigial stamens. Medial longitudinal section of the androecial hood of Lecythis tuyrana. Photo by C. Galdames (SCZ).
Androecial hood: Lecythis persistens subsp. persistens-type The androecial hood of this type is flat and the appendages of the hood are with anthers that bear fodder pollen and, thus, are called staminodes Androecial hood: Lecythis persistens subsp. persistens-type. Photos by S. A. Mori.
Androecial hood: Lecythis pisonis-type The androecial hood is separated from the staminal ring, the hood is flat, and there are staminodes proximally and vestigial stamens distally. Androecial hood: Lecythis pisonis-type. Drawing by Bobbi Angell.
Androecium (pl. = androecia) The male part of the flower. Actinomorphic flowers of Gustavia. Photo by R. Foster.
Androecium actinomorphic Capable of being divided in more than one plane into two or more equal halves; usually, but not always, used to describe flowers. The major types of actinomorphic and zygomorphic flowers of Neotropical Lecythidaceae. Drawings by B. Angell.
Androecium zygomorphic The prolongation of one side of the androecium of all species of , , , , , , and . Staminal ring zygomorphic. Drawing by B. Angell.
Androphore A stalk elevated above the point of perianth attachment which supports the stamens. Androecium of Gustavia johnclarkii. Photo by J. L. Clark.
Anisocytic stomata Referring to a stomatal type in which one of the subsidiary cells surrounding a stomate is smaller than the other epidermal cells. Two stomatal types in Gustavia. Drawing by C. A. Gracie.
Anomocytic stomata Referring to a stomatal type in which the subsidiary cells surrounding a stomate are not differentiated from the other epidermal cells. Two stomatal types in Gustavia. Drawing by C. A. Gracie.
Anterior Two stomatal types in Gustavia. Drawing by C. A. Gracie.
Anterior end of flower The most distal end of the flower but not including inner coils, i.e., the end of a zygomorphic-flowered Lecythidaceae into which a pollinator enters to reap a pollinator reward. Incipient ligular sulcus of Eschweira micrantha. Photo by C. A. Gracie.
Anterior hood extension An extension of the ligule beyond the point where the androecial hood splits into two extensions, this extension is the uppermost division of the androecial hood split and extends forward from the split. Medial sections sections of an androecium of Bertholletia excelsa. Photo by N. P. Smith.
Anther The pollen-bearing part of the stamen. Lateral and poricidal anther dehiscence. Drawing by B. Angell.
Anther connective The tissue that connects the thecae on one side of an anther to the thecae on the other side. Stamens of Grias peruviana. Photo by C. Carollo.
Anther dehiscence lateral Anthers that dehisce by lateral slits. Androecium of Lecythis holcogyne. Photo by N. P. Smith.
Anther dehiscence poricidal Anthers that open (dehisce) and release pollen via a terminal pore. Poricidal anthers and unidimensional filaments. Drawing by B. Antell.
Anther theca (pl. = thecae) The chambers of an anther in which the pollen is produced. Same as pollen sac. Stamens of Grias peruviana. Photo by C. Carollo.
Antherode The anther of a staminode; an anther that bears pollen that does not germinate. Heteranthery in Corythophora. Photos by S. A. Mori.
Anthers basifixed Anthers attached to their filaments at the very base. Filament stalk of Gustavia grandibracteata. Photo by N. P. Smith.
Anthers black Anthers that look black at anthesis either because the anther is black or because the pollen is black. Medial section of the androecium of Eschweilera micrantha. Photo by C. A. Gracie.
Anthers orange Anthers that look orange at anthesis either because the anther is orange or because the pollen is orange. Apical view of a flower of Couroupita nicaraguarensis. Photo by C. Persson.
Anthers versatile Anthers that are attached above the base and can be moved more freely on the apices of their filaments than can anthers that are attached at their bases (= anthers basifixed). Stamens of Grias peruviana. Photo by C. Carollo.
Anthers white Anthers that look white at anthesis either because the anther is white or because the pollen is white. White anthers are common in neotropical Lecythidaceae. Fertile and fodder pollen of Couroupita guianensis. Photo by L. E. Gámez Alvarez.
Anthers yellow Anthers that look yellow at anthesis either because the anther is yellow or because the pollen is yellow. Flower of Lecythis barnebyi showing hood and staminal ring of the zygomorphic androecium. Photo by C. A. Gracie.
Anthesis The expanding and opening of a flower; another way of saying "in flower." Flower of Gustavia augusta with trigonid bees. Photo by S. A. Mori.
Appendage-free ligule The part of a ligule between the staminal ring and the androecial hood that is free of stamens, staminodes, and vestigial stamens (collectively called appendages), or at least the appendages are very sparse compared to their density in the staminal ring or on the hood. Terms associated with a staminal ring zygomorphic-flowered Lecythidaceae with androecial hoods. Drawing by B. Angell.
Appendages Same as androecial appendages. Terms associated with a staminal ring zygomorphic-flowered Lecythidaceae with androecial hoods. Drawing by B. Angell.
Arched tertiary veins Referring to tertiary veins that orginate from the midrib and arch downward to the infra adjacent secondary vein. Leaf venation of Gustavia paucisperma. Photo by S. A. Mori.
Areole (areolate) Referring to the smallest area of leaf tissue surrounded by veins, the sides of which can be comprised of any order of vein (Ellis et al., 2009). Leaf venation of a species of Eschweilera. Photo by S. A. Mori.
Aril A fleshy outgrowth from the base of the funicle. Seeds of Lecythis pisonis.  Photo by C. Gracie.
Aril basal An aril located at the base of the seed. Basal aril of Lecythis pisonis. Photo by S. A. Mori.
Aril basal-lateral An aril starting at the base and extending part way up the side of the seed. Basal-lateral aril of Lecythis mesophylla. Photo by R. Aguilar.
Aril half I-beam An aril that runs the length of the side of the seed and then abruptly turns to cover one or both ends of the seed. Fruit and seeds of Eschweilera antioquensis. Photo by S. A. Mori.
Aril lateral An aril that runs along the side of the seed such as in many, but not all, species of . Lateral aril of Eschweilera. Photo by S. A. Mori
Aril spreading The aril spreads around the entire seed and, thus, looks like a sarcrotesta. In contrast to a sarcotesta, a spreading aril arises from the I-beam aril and not the seed surface. Fresh fruit, seeds, and spreading aril of Eschweilera jefensis. Photo by M. Correa (PMA & SCZ)