Mickel, John T. & Smith, Alan R. 2004. The pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1-1054.
Thelypteridaceae
Species Description - Rhizomes erect; fronds 50 to often 100(–150) cm or more long; stipes stramineous to tan, sometimes mucilaginous when young, mostly 5–20 cm x 3–7 mm diam.; blades to 130 x 40 cm, with to 10 pairs of gradually to subabruptly reduced, widely spaced pinnae proximally; pinnae to (4–)10–16(–22) x 1.5–3 cm, deeply incised to within 1 mm of costae, usually opposite, horizontal, most pinnae with proximal segments somewhat elongate; aerophores peg-like at pinna bases; segments perpendicular or nearly so to costae, linear, 2–3 mm wide; veins to 18 pairs per segment; indument abaxially of sparse to dense hairs on rachises, costae, and costules, hairs unicellular or often septate, to 1.5 mm long, occasionally hairs absent, tissue between veins often with numerous orangish to reddish, sessile, hemispherical glands; sori medial to submarginal, with indusia persistent, thin, tan, often glandular; sporangia glabrous.
Polypodium balbisii Spreng., Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat. Cur. 10: 228. 1821. Aspidium sprengelii Kaulf., Flora 6: 365. 1823, nom. superfl. (see Morton, Amer. Fern J. 53: 62. 1963). Dryopteris sprengelii (Kaulf.) Kuntze, Revis. Gen. Pl. 2: 813. 1891. Dryopteris balbisii (Spreng.) Urb., Symb. Antill. 4: 14. 1903. Thelypteris sprengelii (Kaulf.) Proctor, Bull. Inst. Jamaica Jamaica, Sci. Ser. 5: 65. 1953. Neotype (chosen by Proctor, 1977: 281). Dominica. Hodge & Hodge 1203 (GH, not found, 2002). Complete synonymy given in Smith (1983).
Thelypteris balbisii is closely related to T. resinifera, and it would not be surprising to find both species growing together and hybridizing. Two collections vary in the direction of T. resinifera: Mickel 1301 (NY, UC) and Hallberg 1319 (NY, UC). Usually T. balbisii can be distinguished by the presence of a least a few long, septate hairs on the costae abaxially, by the more spreading and longer pinna segments, by its larger size, by the lack of hairs on the indusia, and by the more remote and fewer reduced proximal pinnae. See T. struthiopteroides for additional comments.
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