Mickel, John T. & Smith, Alan R. 2004. The pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1-1054.
Dryopteridaceae
Species Description - Rhizomes suberect, woody, stout; stipes stramineous to brownish, to 50+ cm x 7 mm, ca. 1/3 the frond length, glabrescent or with scattered, linear, fibrillose scales distally; stipe base scales dense, castaneous, lustrous, linear, 10–20(–30) x 0.1–0.3 mm, with prominent marginal falcate (hooked) teeth, the cells elongate; blades deltate, to 1(–1.5) m long, 3-pinnate-pinnatifid at bases, proximal pinna pair to 1 m long, much more developed basiscopically than acroscopically; middle pinnae mostly 2- pinnate-pinnatifid, sessile to stalked to ca. 1 cm; indument abaxially on rachises, costae, and midveins, of scattered hair-like toothed scales (0.5–)1–4 mm long, sometimes also with sparse hyaline acicular hairs 0.1–0.3 mm long on costae and costules, and sometimes even a few short hairs on laminar tissue between veins; indument adaxially of stout, curved, acicular, septate hairs 0.3–0.8 mm long on costae and costules, veins and tissue between veins usually glabrous; sori supramedial, exindusiate; 2n=ca. 164 (Jam).
Polypodium subincisum Willd., Sp. Pl., ed. 4, 5(1): 202. 1810. Dryopteris subincisa (Willd.) Urb., Symb. Antill. 4: 19. 1903. Phegopteris subincisa (Willd.) Fe´e, Me´m. Foug. 5: 243. 1852. Ctenitis subincisa (Willd.) Ching, Sunyatsenia 5: 250. 1940. Type. Venezuela. Caracas, Bredemeyer s.n. (B-Willd. 19701!). Polypodium galeottii M. Martens, Me´m. Foug. Mexique 43, pl. 7, f. 3. 1842. Type. Mexico. [Veracruz:] “Mirador” [on specimen], Galeotti 6321 (BR!; isotype BR! frag. US!). Phegopteris stenolepis Fe´e, Me´m. Foug. 8: 89. 1857 [“stenolepsis”]. Type. Mexico. Veracruz: Near Huatusco, Schaffner 239 (RB; isotype K!).
Megalastrum subincisum and its close relatives are distinct from Ctenitis species in their toothed rhizome scales and long, often curved, acicular hairs adaxially on the costae and segments; the toothed, narrow scales abaxially on the costae will separate M. subincisum from M. lunense. From M. atrogriseum and M. pulverulentum it differs by the shorter hairs along the costae and costules abaxially. Breedlove 22633 (DS), from Chiapas, is a much scalier variant of M. subincisum; the ultimate segments are also broader and shorter than typical.
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