Monographs Details:
Authority:
Cuatrecasas, José. 1970. Brunelliaceae. Fl. Neotrop. Monogr. 2: 1-189. (Published by NYBG Press)
Cuatrecasas, José. 1970. Brunelliaceae. Fl. Neotrop. Monogr. 2: 1-189. (Published by NYBG Press)
Family:
Brunelliaceae
Brunelliaceae
Description:
Description - Tree about 20 m tall with spreading branches, the terminal subterete, cinereous green, lenticellate, and rather densely pubescent, with ascending, appressed trichomes, substrigose. Stipules two to three on each side of the petiole, subulate, strigose, 6-3 mm long, caducous. Leaves opposite, 2-4-5-jugate, axis 9-24 cm long, the interjuga 4-6 cm long, subterete, very slightly striate and narrowly sulcate above, pubescent with copious, weak, mostly subappressed hairs, the stipels usually bigeminate, aciculate or subcorniculate, 1.5-3.0 mm long; petiole 6-12 cm long, subterete, pubescent, slightly thickened at the base. Leaflets chartaceous, pale greenish; lateral petiolules 5-10 mm long, pubescent, sulcate above, the terminal ones 10-20 mm long; blades 7-15 cm long, 2.5-5.0 cm wide, elliptic-oblong, sublanceolate, asymmetric at the base with one half rounded the other half somewhat decurrent, attenuate and slightly acuminate at the apex, moderately crenate the crenations about 5-8 mm long with minute, acute, apical teeth; light yellowish-green, shining and glabrous above (except for a few hairs on the midrib) the midrib impressed, the secondary nerves noticeable, pale and thin, the minor veins obsolete; greenish-cinereous or pale ochroleucous beneath, pubescent with short, acute, rather rigid, subascendent or subappressed hairs very copious above the nervation, the costa very prominent, subcarinate, the secondary nerves 18-22 pairs, 3-6 mm apart, prominent, parallel, ascending, with an angle of divergence 45°-55°, curving at the margin, ending in a callous minute tooth, the tertiary and quaternary veins prominulously reticulate, the inter-reticular surface strongly papillose, rather cinereous-whitish, contrasting with the more yellowish, pubescent, and sparsely glandulose-punctate venation. Inflorescences axillary, elongate, somewhat shorter than the leaves, with rather long, spreading, loose branching to 30 cm long, the peduncle 8-12 cm long, angulate, compressed, pubescent or puberulous, with ascendent hairs, the branches thin, flexuous, dichotomous, compressed, ancipital, pubescent, and final divisions trichotomous, slender, spreading-pilose. Pedicels pedunculate; bracts linear, pubescent, 0.4-1.0 mm long; peduncles 0.5-2.0 mm long; pedicels 1-2 mm long, pubescent. Buds tetragonous-globose, about 1.8 mm broad, 1.5 mm tall, white, or roseate when dry. Male flowers tetramerous; expanded calyx about 4 mm in diam, the lobes ovate, subacute, about 1.5 mm long, 1.3-1.4 mm wide, with a gland at the sinus, sparsely appressed-pilose outside, densely sericeous inside, minutely tomentulose at the margin; stamens eight, the filaments 1 mm long (in the bud), with a few hairs at the base; anthers thick, ellipsoid, obtuse, 0.8-0.9 mm long; carpels four, rudimentary, about 0.9 mm long, the ovary hyaline-pilose, 0.3 mm long with two rudimentary ovules, the style subulate, almost glabrous, smooth, 0.4-0.5 mm long; disc hirsutulous. Female flowers in fruit also tetramerous; calyx when expanded 5 mm in diameter, the lobes rounded-ovate, 1.8 × 1.8 mm, with a punctiform gland at the sinus; staminodes eight, the filament 1 mm long, glabrous, the anther sterile, oblong, about 0.7 mm long; carpels four, often all developed into fruit; ovary biovulate, ovoid, slightly compressed, sericeous-hirsute, about 1 mm long; style about 2 mm long, the end contorted, the base thickened and minutely sericeous. Follicles 2.5 × 2.5 mm, subglobose, apiculate and slightly compressed, minutely sericeous-tomentose and hispid (the bristles 1 mm long), the style about 2.5 mm long, the base forming appressed-pilose apiculum. Endocarp monospermous, ca 2.3 × 2.3 mm, papyraceous, subglobose with angular insertion point, for the rest smooth, when dry open U-shaped. Seeds subglobose compressed, about 1.7 × 1.3 mm.
Description - Tree about 20 m tall with spreading branches, the terminal subterete, cinereous green, lenticellate, and rather densely pubescent, with ascending, appressed trichomes, substrigose. Stipules two to three on each side of the petiole, subulate, strigose, 6-3 mm long, caducous. Leaves opposite, 2-4-5-jugate, axis 9-24 cm long, the interjuga 4-6 cm long, subterete, very slightly striate and narrowly sulcate above, pubescent with copious, weak, mostly subappressed hairs, the stipels usually bigeminate, aciculate or subcorniculate, 1.5-3.0 mm long; petiole 6-12 cm long, subterete, pubescent, slightly thickened at the base. Leaflets chartaceous, pale greenish; lateral petiolules 5-10 mm long, pubescent, sulcate above, the terminal ones 10-20 mm long; blades 7-15 cm long, 2.5-5.0 cm wide, elliptic-oblong, sublanceolate, asymmetric at the base with one half rounded the other half somewhat decurrent, attenuate and slightly acuminate at the apex, moderately crenate the crenations about 5-8 mm long with minute, acute, apical teeth; light yellowish-green, shining and glabrous above (except for a few hairs on the midrib) the midrib impressed, the secondary nerves noticeable, pale and thin, the minor veins obsolete; greenish-cinereous or pale ochroleucous beneath, pubescent with short, acute, rather rigid, subascendent or subappressed hairs very copious above the nervation, the costa very prominent, subcarinate, the secondary nerves 18-22 pairs, 3-6 mm apart, prominent, parallel, ascending, with an angle of divergence 45°-55°, curving at the margin, ending in a callous minute tooth, the tertiary and quaternary veins prominulously reticulate, the inter-reticular surface strongly papillose, rather cinereous-whitish, contrasting with the more yellowish, pubescent, and sparsely glandulose-punctate venation. Inflorescences axillary, elongate, somewhat shorter than the leaves, with rather long, spreading, loose branching to 30 cm long, the peduncle 8-12 cm long, angulate, compressed, pubescent or puberulous, with ascendent hairs, the branches thin, flexuous, dichotomous, compressed, ancipital, pubescent, and final divisions trichotomous, slender, spreading-pilose. Pedicels pedunculate; bracts linear, pubescent, 0.4-1.0 mm long; peduncles 0.5-2.0 mm long; pedicels 1-2 mm long, pubescent. Buds tetragonous-globose, about 1.8 mm broad, 1.5 mm tall, white, or roseate when dry. Male flowers tetramerous; expanded calyx about 4 mm in diam, the lobes ovate, subacute, about 1.5 mm long, 1.3-1.4 mm wide, with a gland at the sinus, sparsely appressed-pilose outside, densely sericeous inside, minutely tomentulose at the margin; stamens eight, the filaments 1 mm long (in the bud), with a few hairs at the base; anthers thick, ellipsoid, obtuse, 0.8-0.9 mm long; carpels four, rudimentary, about 0.9 mm long, the ovary hyaline-pilose, 0.3 mm long with two rudimentary ovules, the style subulate, almost glabrous, smooth, 0.4-0.5 mm long; disc hirsutulous. Female flowers in fruit also tetramerous; calyx when expanded 5 mm in diameter, the lobes rounded-ovate, 1.8 × 1.8 mm, with a punctiform gland at the sinus; staminodes eight, the filament 1 mm long, glabrous, the anther sterile, oblong, about 0.7 mm long; carpels four, often all developed into fruit; ovary biovulate, ovoid, slightly compressed, sericeous-hirsute, about 1 mm long; style about 2 mm long, the end contorted, the base thickened and minutely sericeous. Follicles 2.5 × 2.5 mm, subglobose, apiculate and slightly compressed, minutely sericeous-tomentose and hispid (the bristles 1 mm long), the style about 2.5 mm long, the base forming appressed-pilose apiculum. Endocarp monospermous, ca 2.3 × 2.3 mm, papyraceous, subglobose with angular insertion point, for the rest smooth, when dry open U-shaped. Seeds subglobose compressed, about 1.7 × 1.3 mm.
Distribution:
Ecuador South America| Pichincha Ecuador South America|
Ecuador South America| Pichincha Ecuador South America|