Monographs Details:
Authority:
Cuatrecasas, José. 1970. Brunelliaceae. Fl. Neotrop. Monogr. 2: 1-189. (Published by NYBG Press)
Cuatrecasas, José. 1970. Brunelliaceae. Fl. Neotrop. Monogr. 2: 1-189. (Published by NYBG Press)
Family:
Brunelliaceae
Brunelliaceae
Description:
Description - Tree 25 m tall, the terminal branchlets angulate-sulcate, compressed, puberulous, soon glabrous. Terminal buds pubescent with ascendent, appressed or subappressed hairs. Stipules subulate, pubescent, single on each side of the petiole, 1.0-1.5 mm long. Leaves opposite, 4-9-jugate; axis 9-24 cm long, shortly and rather spreading-pubescent, the interjuga 1.2-3.5 cm long, subterete, minutely striolate, narrowly sulcate above; stipels geminate, subulate, pubescent, 1 mm long, soon caducous; petiole subterete, slightly thickened towards the base, 4.5-12.0 cm long. Leaflets opposite, rigidly chartaceous; petiolules 1-4 mm long, the terminal ones 5-18 mm long; blades 4.5-14.5 cm long, 2-4 cm wide, elliptic-oblong, obtuse and asymmetric at the base, attenuate and acuminate at the apex, the acute acumen often subhamate, 1.0-1.5 cm long, serrulate-crenate, the teeth 5-10 mm apart, accumbent, callous, acute; above green, rather nitid, and glabrous except for the appressed-puberulous midrib and a few sparse hairs, the costa impressed, the secondary nerves and minor veins somewhat prominulous and conspicuous; cinereous-green and pubescent beneath, the costa prominent, terete or striolate and carinate, the secondary nerves in 17-22 pairs, 4-6 mm apart, parallel, prominent, slightly ascending with an angle of divergence of 50°-65°, curving and anastomosing at the margin, the tertiary nerves forming a lax prominulous reticulum with other veins, the smaller veins forming a minor minute reticulum flat at the bottom, the pubescence rather angular-ascending or of subappressed hairs on the main nervation and small spreading hairs on the reticulum, the intervenal surface densely whitish-papillose. Panicles axillary, 4.5-20.0 cm long and broad but usually broader than long, the peduncle 1-5 cm long, stout, straight, compressed, striate, pubescent with patulous small hairs, the first branching trichotomous, the rest regularly dichotomous, the branchlets rather short, angular, spreading-pubescent, the ultimate very short, bearing the shortly pedicellate, glomerate flowers. Bracts subulate, 0.2-1.0 mm long, deciduous. Pedicels sessile or short stalked, 0.2-1.0 mm long, pubescent, with ascendent or patulous hairs. Buds globose, about 2 mm in diam. Flowers pentamerous or tetramerous. Male flowers: calyx 2.2-2.5 mm long, when expanded 5 mm in diam, the lobes ovate, acute, 0.8-2.1 mm long, 1.0-1.2 mm wide, pubescent with appressed, ascendent hairs outside, villous on the nerves inside, the margin minutely tomentulose; fertile stamens eight or more often ten, the filaments about 4 mm long with hyaline, patulous hairs on the lower part; anthers ellipsoid, obtuse, 1.0-1.1 mm long; sterile carpels rudimentary with minute, obsolete hirsutulous ovary and capillary, glabrous, style, 1 mm long. Female flowers (in immature fruit), with expanded calyx 4.5 mm in diam, the lobes about 1.6 mm long, 1.2 mm broad; staminodia ten or eight, the filaments glabrous, 1 mm long, the sterile anthers oblong, 0.6 mm long; ovaries ellipsoid-ovoid, 1.0-1.2 mm long, tomentose and hispidulous, the style 2 mm long. Disc hirsute-tomentose. Follicles ellipsoid-ovoid, 2.5-2.8 × 2.0-2.2 mm, sericeous and hispid; endocarp cartilaginous, still soft in immature specimens, 2.2 × 2 mm, monospermous. Seeds ellipsoid, compressed, about 2.0 × 1.3 mm.
Description - Tree 25 m tall, the terminal branchlets angulate-sulcate, compressed, puberulous, soon glabrous. Terminal buds pubescent with ascendent, appressed or subappressed hairs. Stipules subulate, pubescent, single on each side of the petiole, 1.0-1.5 mm long. Leaves opposite, 4-9-jugate; axis 9-24 cm long, shortly and rather spreading-pubescent, the interjuga 1.2-3.5 cm long, subterete, minutely striolate, narrowly sulcate above; stipels geminate, subulate, pubescent, 1 mm long, soon caducous; petiole subterete, slightly thickened towards the base, 4.5-12.0 cm long. Leaflets opposite, rigidly chartaceous; petiolules 1-4 mm long, the terminal ones 5-18 mm long; blades 4.5-14.5 cm long, 2-4 cm wide, elliptic-oblong, obtuse and asymmetric at the base, attenuate and acuminate at the apex, the acute acumen often subhamate, 1.0-1.5 cm long, serrulate-crenate, the teeth 5-10 mm apart, accumbent, callous, acute; above green, rather nitid, and glabrous except for the appressed-puberulous midrib and a few sparse hairs, the costa impressed, the secondary nerves and minor veins somewhat prominulous and conspicuous; cinereous-green and pubescent beneath, the costa prominent, terete or striolate and carinate, the secondary nerves in 17-22 pairs, 4-6 mm apart, parallel, prominent, slightly ascending with an angle of divergence of 50°-65°, curving and anastomosing at the margin, the tertiary nerves forming a lax prominulous reticulum with other veins, the smaller veins forming a minor minute reticulum flat at the bottom, the pubescence rather angular-ascending or of subappressed hairs on the main nervation and small spreading hairs on the reticulum, the intervenal surface densely whitish-papillose. Panicles axillary, 4.5-20.0 cm long and broad but usually broader than long, the peduncle 1-5 cm long, stout, straight, compressed, striate, pubescent with patulous small hairs, the first branching trichotomous, the rest regularly dichotomous, the branchlets rather short, angular, spreading-pubescent, the ultimate very short, bearing the shortly pedicellate, glomerate flowers. Bracts subulate, 0.2-1.0 mm long, deciduous. Pedicels sessile or short stalked, 0.2-1.0 mm long, pubescent, with ascendent or patulous hairs. Buds globose, about 2 mm in diam. Flowers pentamerous or tetramerous. Male flowers: calyx 2.2-2.5 mm long, when expanded 5 mm in diam, the lobes ovate, acute, 0.8-2.1 mm long, 1.0-1.2 mm wide, pubescent with appressed, ascendent hairs outside, villous on the nerves inside, the margin minutely tomentulose; fertile stamens eight or more often ten, the filaments about 4 mm long with hyaline, patulous hairs on the lower part; anthers ellipsoid, obtuse, 1.0-1.1 mm long; sterile carpels rudimentary with minute, obsolete hirsutulous ovary and capillary, glabrous, style, 1 mm long. Female flowers (in immature fruit), with expanded calyx 4.5 mm in diam, the lobes about 1.6 mm long, 1.2 mm broad; staminodia ten or eight, the filaments glabrous, 1 mm long, the sterile anthers oblong, 0.6 mm long; ovaries ellipsoid-ovoid, 1.0-1.2 mm long, tomentose and hispidulous, the style 2 mm long. Disc hirsute-tomentose. Follicles ellipsoid-ovoid, 2.5-2.8 × 2.0-2.2 mm, sericeous and hispid; endocarp cartilaginous, still soft in immature specimens, 2.2 × 2 mm, monospermous. Seeds ellipsoid, compressed, about 2.0 × 1.3 mm.
Distribution:
Ecuador South America| Napo Ecuador South America| Pastaza Ecuador South America|
Ecuador South America| Napo Ecuador South America| Pastaza Ecuador South America|