Taxon Details: Arbutus xalapensis (Benth.) Kunth
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Narratives:

Family:

Ericaceae (Magnoliophyta)
Scientific Name:

Arbutus xalapensis (Benth.) Kunth
Primary Citation:

Nov. Gen. Sp. (Kunth) 3: 279. 1819
Accepted Name:

This name is currently accepted.
Common Names:

encino roble, flora de pulca, guayavillo, indio desnudo, korúvasi, kurúvasi, madrón, madrona, madrone, madroña, madroño colorado, nuzundu, pách-sich-ách-mixe, urúbasi, ya-hatzii, madroño
Description:

Description: Trees or arborescent shrubs, 4-6 m, often up to 15 m (occasionally said to be taller); bark usually brick red and peeling in large, smooth flakes over most of the larger limbs, ceasing to exfoliate at the base of the trunk on older specimens and eventually retained over most of the oldest parts of the plant, becoming gray, irregularly roughened; twigs of newly emerging shoots usually densely villous, often with a mixture of glandular hairs, fully developed twigs pubescent and/or glandular pubescent, or glabrate to completely glabrous, twig bark soon loosening and exfoliating, older twigs usually smooth, brick-red, or glaucous grayish-red. Leaves pale or bright olive-green or glaucous-green, slightly lighter beneath, blades elliptic or slightly ovate-elliptic, (4-)5-11(-15) x 1.5-4.8(-6) cm, basally tapered, rounded, sub-truncate, or slightly cordate, apically acute or obtuse, the margins smooth or irregularly toothed, rarely slightly spinulose, upper surface glossy-glabrous (when fresh) or pubescent, often densely so toward base of blade and especially along the midrib, lower surface glabrous or more often pubescent, frequently white, tan, or brown woolly, sometimes with a few glandular hairs; petiole 1/4-1/3(-1/2) the length of the blade, vestiture about the same as on the blade, but glandular hairs more dense if present, these straight or curly, drying stiff, up to 1.5 mm long. Inflorescence a terminal cluster of racemes, very variable, from densely clustered or openly-branched, to few-flowered, often showy, axes including pedicels usually with glandular hairs, these sometimes copious and conspicuous. Flowers borne obliquely erect on accrescent pedicels 6-9(-14) mm long, subtended by a reddish or tan-colored accrescent bract 2-3.5 mm long, to 4.9 mm long in fruit, enclosing 2 smaller bracteoles; calyx 1.8-3 mm long (northern material with a blush of pink), the lobes often dorsally pubescent, margins scarious and ciliate or glandular-ciliate; corolla 5.1-6(-7.2) mm long, the larger on rapidly developing, more open and elongated inflorescences, inner surface of tube smooth or sparsely pubescent below the middle; filaments 2.1-3 mm long; anthers about 1.5 mm long, spurs (1/2-)3/4-4/5 the length of the thecae; ovary with up to 10 ovules per locule. Fruit more or less spherical, or slightly turbinate, 7.5-8.8(-9) mm diam. when ripe; seeds 4 or 5 per locule, 1.8-2.5 mm long; chromosome number: 2n=26 (Callan, 1941).

Distribution: Southern New Mexico and W Texas, U.S.A., south in mountainous topography through all the states of Mexico (except Campeche, Quintana Roo, Tabasco, and Yucatán), into Guatemala, Honduras, El Salvador, and Nicaragua, at elevations of (325-)2000-3000(-3400) m, at highest elevation in the Sierra Volcánica Transversal crossing southern Mexico. Flowering in Central America and southeastern Mexico November to mid-February, progressively later into April and May in the northernmost localities, rarely during mid-year; fruiting follows after about six weeks.

Local names: Throughout its range Arbutus xalapensis is called madrone (English), madroño (Spanish), less frequently madroña (Spanish), or simply madrón (Spanish). Additional vernacular names: encino roble, flora de pulca, guayavillo, indio desnudo, kurúvasi or korúvasi, madroño colorado, nuzu-ndu, pách-sich-ách-mixe, urúbasi, and ya-hatzii.

Cultivated: E, MO.

Type: Mexico. Veracruz: Near Xalapa, 1260 m, Feb 1804 (fl), Humboldt & Bonpland s.n. (lectotype: designated by Sørensen, 1995, P; Microfiche, IDC 6209, fiche no. 78).

Flora and Monograph Treatment(s):

Arbutus xalapensis (Benth.) Kunth: [Article] Luteyn, James L., et al. 1995. Ericaceae, Part II. The Superior-Ovaried Genera (Monotropoideae, Pyroloideae, Rhododendroideae, and Vaccinioideae P.P.). Fl. Neotrop. Monogr. 66: 560.
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P. Stanley 81496, Guatemala
M. P. Martinez 62, Mexico
C. H. T. Townsend 208, Mexico
C. G. Pringle 1322, Mexico
C. G. Pringle 1322, Mexico
C. V. Hartman 587, Mexico
208, Mexico
D. S. Correll 21774, Mexico
F. Chiang Cabrera 10058, Mexico
F. Chiang Cabrera 10127, Mexico
J. S. Henrickson 15287, Mexico
J. S. Henrickson 14795, Mexico
W. C. Holmes 4496, Mexico
11146, Mexico
9928, Mexico
P. Tenorio L. 715, Mexico
R. D. Worthington 7891, Mexico
D. S. Correll 20116, Mexico
E. W. Nelson 5003, Mexico
R. W. Spellenberg 10043, Mexico
C. G. Pringle 1321, Mexico
J. Spencer 585, Mexico
D. J. Rosen 4470, Mexico
R. Hernández R. 136, Mexico
C. G. Pringle 1322, Mexico
D. S. Correll 21491, Mexico
D. S. Correll 21485, Mexico
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G. M. Diggs, Magnoliophyta; North America
C. G. Pringle 8002, Mexico
G. M. Diggs, Magnoliophyta; North America
D. J. Pinkava 6093, Mexico
M. C. Johnston 10945, Mexico
M. C. Johnston 10155, Mexico
E. Joyal 1649, Mexico
8, Mexico
L. M. González Villarreal 2861, Mexico
H. H. Iltis 7822B, Mexico
T. S. Cochrane 10635a, Mexico
G. M. Diggs, Magnoliophyta; North America
R. D. Worthington 7932, Mexico
H. S. Gentry 5878, Mexico
E. Palmer 578, Mexico
E. J. Lott 2060, Mexico
P. D. Sørensen 7936, Mexico
C. G. Pringle 2444, Mexico
C. G. Pringle 2444, Mexico
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598, Mexico
7012, Mexico
C. G. Pringle 8033, Mexico
C. G. Pringle 11007, Mexico
C. G. Pringle 6814, Mexico
C. G. Pringle 6813, Mexico
J. N. Rose 8480, Mexico
C. A. Purpus 1773, Mexico
J. G. Schaffner 314, Mexico
6152, Mexico
7351, Mexico
C. E. Smith, Magnoliophyta; North America
3519, Mexico
G. M. Diggs, Magnoliophyta; North America
G. D. McPherson 897, Mexico
646, Mexico
C. G. Pringle 6846, Mexico
C. G. Pringle 8011, Mexico
G. M. Diggs, Magnoliophyta; North America
E. K. Balls 4874, Mexico
D. E. Breedlove 58979, Mexico
G. J. Breckon 1282, Mexico
G. M. Diggs, Magnoliophyta; North America
R. C. Rollins 5858, Mexico
E. M. Chittenden 117, Mexico
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J. Espinosa G. 2055, Mexico
J. Dorantes 140, Mexico
W. Marquez R. 38W, Mexico
G. F. Ledingham 4618, United States of America
E. K. Balls B4398, Mexico
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C. A. Purpus 2994, Mexico
A. Lot Helgueras 1066, Mexico
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X. Munn-Estrada 1689, Mexico
J. Pérez 376, Mexico
J. J. Castillo 2384, Guatemala
V. E. Grant 681, Guatemala
P. R. House 1156, Honduras
H. Thomas 294, Honduras
L. Morales R. 141, Honduras
J. M. Tucker 1108, El Salvador
P. H. Allen 7183, El Salvador
D. E. Breedlove 31457, Mexico
J. Conrad 2079, Mexico
D. E. Stone 2788, Mexico
R. Torres Colín 1327, Mexico
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H. G. Galeotti 1844, Mexico
H. G. Galeotti 1843, Mexico
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E. Contreras 4884, Guatemala
C. L. Smith 202, Mexico
C. G. Pringle 6110, Mexico
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P. Stanley 66456, Guatemala
W. A. Kellerman 7236, Guatemala
R. W. Spellenberg 8425, Mexico
P. Tenorio L. 1891, Mexico
R. W. Spellenberg 8481, Mexico
E. Palmer 325, Mexico
S. Gonzalez 2661, Mexico
E. Palmer 68, Mexico
V. W. Steinmann 5329, Mexico
A. Ventura A. 475, Mexico
J. A. Pérez de la Rosa 3060, Mexico
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B. M. Bartholomew 2681, Mexico
377, Mexico
G. L. Fisher 35439, Mexico
C. A. Purpus 2995, Mexico
C. G. Pringle 1886, Mexico
C. G. Pringle 1108, Mexico
C. G. Pringle 10255, Mexico
C. G. Pringle 1886, Mexico
G. Arsène 5354, Mexico
W. C. Leavenworth 569, Mexico
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S. D. Koch 77373, Mexico
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R. Hernández Magaña 1030, Mexico
E. K. Balls B5454, Mexico
E. Matuda 1227, Mexico
J. Dorantes 5122, Mexico
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R. M. Laughlin 637, Mexico
A. S. Ton 1818, Mexico
J. I. Calzada 19661, Mexico
A. S. Ton 9447, Mexico
M. López 451, Mexico
A. S. Ton 417, Mexico
L. M. López López 0066, Mexico
P. Tenorio L. 2024, Mexico
D. E. Breedlove 31183, Mexico
C. P. Cowan 4030, Mexico
R. E. Gereau 1972, Mexico
L. Hernández Garcia 0191, Mexico
N. Maldonado Vásquez 0120, Mexico
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F. L. Wynd 505, Mexico
R. Hernández R. 333, Mexico
C. G. Pringle 7367, Mexico
C. Robles P. 168, Mexico
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J. I. Calzada 22547, Mexico
S. Avendaño Reyes 88, Mexico
J. I. Calzada 2099, Mexico
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Y. E. J. Mexia 2500, Mexico
Y. E. J. Mexia 847, Mexico
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K. Velasco Gutiérrez 2000, Mexico
W. H. Camp 2308, Mexico
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J. Renfrow 150, Honduras
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R. A. Bye 9150a, Mexico
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R. A. Bye 9132, Mexico
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Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
Arbutus xalapensis (Benth.) Kunth
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R. Ortega O. 111, Mexico
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