Astragalus umbraticus
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Title
Astragalus umbraticus
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Authors
Rupert C. Barneby
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Scientific Name
Astragalus umbraticus E.Sheld.
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Description
122. Astragalus umbraticus
Rather tall but often quite slender, appearing nearly glabrous but bearing a few scattered, straight, appressed or narrowly ascending hairs up to 0.3-0.7 mm. long on the stems distally and on the margins and midribs of the deep or pale green leaflets, the inflorescence thinly black-strigulose; stems several, diffuse or ascending from the knotty root-crown or shortly forking caudex, (2) 2.5-5 dm. long, pale green becoming stramineous, striate, simple or nearly so, stipules 3.5-9.5 mm. long, the lowest early becoming papery and brownish, embracing 3/4 to nearly the whole stem’s circumference, the upper ones submembranous, longer and narrower, triangular or lance-acuminate, thinly ciliate; leaves 4-12 cm. long, the uppermost shortly petioled or subsessile, with (11) 15-23 broadly oblong, ovate, or obovate, or (in some lower leaves) suborbicular, retuse or abruptly short-acuminate, flat, thin-textured leaflets (4) 6-16 (20) mm. long; peduncles erect or incurved-ascending, 5-12 cm. long, ± equaling the leaf; racemes loosely but shortly 10-25- flowered, the flowers early horizontal, at length declined, the axis little elongating, (1.5) 2-5 cm. long in fruit; bracts membranous, narrowly lanceolate, (1.7) 2-4 mm. long; pedicels at anthesis ascending or a little arched outward, 0.6-1.2 mm. long, in fruit ascending or more strongly arched, a little thickened, scarcely longer; bracteoles 0; calyx 5.2-7 mm. long, strigulose or pilosulous with dark hairs, the somewhat oblique disc 0.7-1.2 mm. deep, the campanulate tube 3.1-4 mm. long, 2.1-3.2 mm. in diameter, the subulate or lance-subulate teeth 2.1-3.2 mm. long; petals greenish-white, often drying cream-color, immaculate; banner recurved through 40-60°, narrowly to broadly ovate-cuneate or rhombic-elliptic, 10-14 mm. long, 5-8 mm. wide; wings 8.4-11 mm. long, the claws 3.5-4.8 mm., the claws 3.5-4.8 mm., the oblong-oblanceolate, obtuse, obliquely truncate and erose, or shallowly emarginate, slightly incurved blades 6-7.6 mm. long, 2-2.9 mm. wide; keel 7.7-10 mm. long, the claws 3.7-5.6 mm., the half-obovate or lunately half-elliptic blades 3.8-5 mm. long, 2.1—3 mm. wide, incurved through 95-100° to the bluntly deltoid apex; anthers (0.35) 0.4—0.55 mm. long; pod spreading or declined, stipitate but the obscure stipe only 0.8-1.9 mm. long, concealed by the calyx, the body lunately or falcately linear-lanceolate or linear in profile, evenly incurved through ¼-½ circle, 1.4—2.4 cm. long, 2.6—3.6 mm. in diameter, cuneate at base, acuminate and cuspidate at apex, laterally compressed- triquetrous, with low-convex lateral and narrower, narrowly but deeply grooved dorsal faces, keeled ventrally by the salient suture, the firmly papery, glabrous valves at first green becoming stramineous and ultimately blackish, delicately cross- reticulate, inflexed as a complete septum 1.5—2.3 mm. wide; ovules 10—15; seeds compressed-quadrangular, brown, smooth but dull, 2.2-2.6 mm. long.—Collections: 15 (o); representative: Jones (from Glendale) in 1902 (CAS, NY, POM); C. L. Hitchcock 19,381 (UC, WS); W. H. Baker 858 (ID, RSA); Tracy 4575 (UC); Peck 8209 (WILLU); Quick 57-24 (CAS).
Dry open oak and pine woodlands, ± 600—4000 feet, uncommon, outer Coast Ranges of Oregon and northern California, apparently most frequent in the valleys of the Coquille, Rogue, and Illinois Rivers in southern Douglas and Josephine Counties, extending north, apparently greatly isolated, to Yamhill County, and south in widely scattered stations to the lower Trinity River and Redwood Creek in Humboldt County, California; not known from the Umpqua drainage in the Coast Ranges, but extending east on the headwaters of that stream into the Cascade foothills (20 miles northeast of Tiller, Douglas County, Quick 57—24, CAS).—Map No. 50.—May to July.
Astragalus umbraticus (of shady habitat) Sheld. in Minn. Bot. Stud. 1: 23. 1894, a substitute for Asylvaticus (of woodland) Wats. in Proc. Amer. Acad. 23: 262. 1888 (non Willd., 1801).—"Near Glendale, in southern Oregon, ‘in open gravelly ground’ (L. F. Henderson) or in dense forests (Thomas Howell, June, 1887)—who suggests the name."—Lectotypus, T. Howell 674, collected June 30, 1887, GH! isotypi (some not or differently numbered), ORE, WS, WTU! paratypi, Henderson 1342, GH, ND, ORE ("open ground along the railroad"). —Hamosa umbratica (Wats.) Rydb. in Bull. Torr. Club 54: 19. 1927.
I have no field experience with the sylvan milk-vetch, A. umbraticus, and other collectors have eft almost no record of its ecology. There is presumptive evidence that it occurs, at least sometimes, on metamorphic and serpentine formations. Its center of dispersal lies in the Klamath Highland, a region rich in endemic relics. The homogeneous nature of the material studied, considered in the light of the wide morphological gap between the species and its nearest relatives, is suggestive of age and a long period of isolation. The broad, flat, green and nearly glabrous leaflets, the small, white flowers, and the glabrous, compressed-triquetrous, strongly but evenly incurved pods are distinctive.