Brunellia trianae Cuatrec.
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Authority
Cuatrecasas, José. 1970. Brunelliaceae. Fl. Neotrop. Monogr. 2: 1-189. (Published by NYBG Press)
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Family
Brunelliaceae
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Scientific Name
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Type
Type. J. J. Triana 5971-4, Colombia, Antioquia; "Andes de Antioquia 2400 m alt, Jun 1852," J' flowers (holotype, COL, photo, FM-44790; F, frag, isotype).
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Description
Description - Tree to about 10 m tall; terminal branches obtusely quadrangular, striate, strongly compressed ancipital at the ends, and as with the terminal buds and folded young leaves extremely appressed cinereous-tomentose, the trichomes flexuous, very densely entangled, forming a tiered layer with a squamose appearance; older branches tending to be terete, glabrate, grayish brown, rugose-striate, lenticellate-tuberculate; internodes 1.5-3.0 cm long. Stipules bigeminate, triangular, the proxals 1-2 mm long, the distals 0.5-1.0 mm long, sometimes extremely minute. Leaves simple, opposite, thick and rigidly coriaceous, fragile. Petiole 1.0-2.5 cm long, robust, thinly striate, above sulcate, laterally compressed, dorsally carinate, appressed arachnoid-tomentose. Blade 10-22 cm long, 4-9 cm wide, oblong-elliptic or ovate-oblong, rounded or abruptly and shortly attenuate at the base, rounded or obtuse at the apex, sometimes slightly attenuate and shortly (to 3 mm) and abruptly acuminate, weakly bicrenate-serrate, the teeth extremely short, subacute, almost obsolete; above ochraceous, appressed lanuginose, caducous indumentum when very young, soon glabrous, yellowish green, lustrous, with very thin but prominent costa in the bottom of a medial furrow, the secondary and tertiary nerves impressed and apparent; below cinereous, with very compact and adherent, lanate tomentum, the costa robust, very prominent, finely striate, laterally compressed, carinate, the secondary nerves in 26-30 pairs, prominent, parallel, 4-9 mm apart, subascendent with a divergence angle of 50°-65°, curved and anastomosing at the margin, the tertiary nerves obliquely transverse, prominent, together with the quaternary veins conspicuously reticulate, the venation deciduous-lanuginose, more or less glabrate, the much smaller reticulum of smaller veins with minute but deep alveolae, covered with minute, adpressed, lanate indumentum. Panicles axillary at the terminal branchlets, about the same length as the subtending leaves (8-17 cm long), the peduncle 3-6 cm long ancipital, striate, appressed lanate; main branches mostly trichotomous or dichotomous, also striate and ancipital, the following ascending, dichotomous divisions angulate, shortly lanuginose, rather thin. Bracts deciduous, lanceolate, 3-2 mm long, the fertile bracteoles strictly linear-subulate, 2-1 mm long, three per flower. Pedicels thin, angulate, minutely crisp-tomentose 1-3 mm long. Flowers tetramerous. Male flowers: buds depressed globose, 3 mm broad, 2 mm high; calyx 2.5 mm high, expanded 6 mm in diam, the lobes triangular, acute, 2.0-2.2 mm long, shortly crisp-tomentose outside, at the margin and apex, almost glabrous inside; stamens eight, the filaments 3.0-3.2 mm long, thick in the lower part, capillary towards the apex, somewhat hairy only at the base, the rest glabrous; anthers ellipsoid, attenuate-apiculate at the apex, cordate at the base, 1 mm long; carpels four, rudimentary, sterile; disc densely tomentose. Female flowers: calyx about 6 mm in diam, 2.8-3.0 mm high, the lobes 2.0-2.3 mm long, triangular, acute, crisp-tomentose outside, apex and margins, almost glabrous inside; staminodia eight, the filaments glabrous, thickening near the base, about 0.5 mm long, the anthers elliptical, attenuate, about 0.6 mm long, sterile; carpels four, the ovary about 1.5 mm long, ovoid, densely tomentose, biovulate; style about 2 mm long, thick, the lower part tomentose. One to four follicles developed to maturity in a flower, about 8 mm long, 4 mm broad, obovoid-oblong, apiculate, shortly and densely lanuginose, monospermous or dispermous; immature seeds reddish, elliptical, about 3 mm long. As described, the tetramerous structure of these flowers is typical, and only very rarely was a calyx found with five lobes.
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Distribution
(Fig 62, p 150.) Central Cordillera of Colombia in Antioquia, in Andean forests between 2400 and 3200 m alt.
Colombia South America| Antioquia Colombia South America|