Banisteriopsis alternifolia (Steyerm.) Cuatrec.
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Authority
Gates, Bronwen. 1982. Banisteriopsis, Diplopterys (Malpighiaceae). Fl. Neotrop. Monogr. 30: 1-238. (Published by NYBG Press)
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Family
Malpighiaceae
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Scientific Name
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Type
Type. J. A. Steyermark 61914, Venezuela, Monagas, Cerro de la Cueva de Doha Anita, fl (holotype, F).
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Synonyms
Banisteria alternifolia Steyerm.
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Description
Species Description - Liana, the branches sparsely appressed-sericeous at first, eventually glabrate, stipules minute, triangular. Leaves alternate, coriaceous, with the petiole 8-15 mm long, sparsely appressed-sericeous to glabrate, bearing a pair of sessile glands apically, with the lamina 4.6-10.6 cm long, 2.3-6.4 cm wide, ovate, obtuse to truncate at the base, obtuse to acuminate at the apex, with the margin bearing minute glands apically, adaxially glabrous, abaxially at first sparsely appressed-sericeous, the hairs sessile, with the trabecula 0.2-0.5 mm long, soon glabrate, with the reticulation prominent on both sides. Inflorescence axillary, the branches up to 30 cm long, silvery appressed-sericeous, the hairs with the trabecula 0.150.25 mm long, bearing bract-like leaves up to 5 mm long subtending racemes of 910 flowers borne alternately; bracts and bracteoles subtending the pedicels 0.6-1.2 mm long, acute at the apex, glabrous, involucrate, persistent; pedicels sessile, 5-10 mm long, 0.4-0.6 mm wide. Sepals glabrous on both sides, 2.6-3.0 mm long, 2.0-2.2 mm wide, rounded at the apex, the 4 lateral sepals Eglandular, the glands 1.2-1.3 mm long, 0.7-0.9 mm wide. Petals yellow, laciniate, the 4 lateral petals reflexed between the sepals, with the claw 1.5-2.0 mm long and up to 0.4 mm wide, the limb 5.0-6.0 mm long and 4.5-6.0 mm wide, orbicular, the antero-lateral petals somewhat larger and more concave than the postero-lateral petals, the posterior petal with the claw erect, 3.0 mm long and up to 0.6 mm wide, canaliculate at the apex, the limb up to 4.5 mm long and 4.0 mm wide, broadly obovate, with the basal laciniae gland-tipped. Stamens with the filaments 2.2-3.6 mm long, those opposite the 3 anterior sepals and the postero-lateral petals longest, those opposite the postero-lateral petals almost the same length as those opposite the antero-lateral sepals, those of the 3 posterior stamens inflexed between the posterior styles, the anthers with the locules glabrous, 0.8-1.0 mm long, the connectives of the stamens opposite the 3 anterior sepals 1.0-1.2 mm long, 0.6-0.8 mm wide and 0.6-0.8 mm deep, broadly obovate, overtopping the locules 0.2-0.4 mm, those of the stamens opposite the petals and the posterolateral sepals 0.6-0.9 mm long, 0.4-0.6 mm wide, 0.2-0.6 mm deep, oblong, not overtopping the locules. Ovary 1.0 mm tall, sparsely pubescent, anterior style straight and erect, 2.6-2.8 mm long, the posterior styles diverging, 2.8-3.0 mm long, stigmas capitate. Fruit unknown.
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Discussion
Collected in flower in March and April.
This is the only species in the genus with alternate leaves, and the flowers are also borne alternately. It is possible that this is a developmental abnormality and does not warrant specific recognition, but I prefer for the moment to maintain its specific identity as it is such a unique species in this respect. It is probably most closely related to Banisteriopsis polygama with which it shares the following characteristics: relatively small coriaceous leaves; silvery-appressed pubescence in the inflorescence; connectives of the stamens opposite the three anterior sepals only a little larger than the others; styles subequal. -
Distribution
Known only from two collections, one in Trinidad and one in Monagas, Venezuela.
Venezuela South America| Monagas Venezuela South America| Trinidad and Tobago South America|