Astragalus oniciformis

  • Title

    Astragalus oniciformis

  • Authors

    Rupert C. Barneby

  • Scientific Name

    Astragalus oniciformis Barneby

  • Description

    131. Astragalus oniciformis

    Slender but wiry, diffuse, perennial, with a taproot and at length loosely forking, suffruticulose caudex, loosely strigulose or subvillosulous with incumbent or ascending, nearly straight or sinuous hairs up to 0.35-0.5 (0.6) mm. long, the herbage greenish-cinereous, the leaflets glabrous or medially glabrescent above; stems several or numerous, decumbent or prostrate, (0.5) 1-2.5 dm. long, commonly spurred at the lowest nodes and thence floriferous upward from all succeeding axils; stipules papery-membranous or early becoming so, 1.5-3.5 (4) mm. long, triangular or triangular-acuminate, decurrent around half, or the lowest around the whole stem’s circumference but not connate; leaves 2.5-1.5 cm. long, shortly petioled or the upper ones subsessile, with stiff but slender rachis and (13) 17—25 (27) rather distant, scattered, narrowly to broadly elliptic, oval, or oblong, obtuse or retuse, mostly folded leaflets 1-6.5 mm. long; peduncles 0.5-2.5 cm. long, much shorter than the leaves; racemes very loosely (4) 6-12-flowered, the flowers early nodding, the axis becoming 1-7 cm. long in fruit, usually produced beyond the last flower as a subulate appendage; bracts papery-membranous, ovate or lanceolate, 0.5-1.8 mm. long; pedicels filiform, very early arched outward or ultimately recurved, at anthesis 1-1.4 mm., in fruit 1-1.6 mm. long, tardily disjointing with the fruit; bracteoles 0; calyx 2.9-3.8 mm. long, strigulose-villosulous like the herbage with white or some dark hairs, the subsymmetric disc 0.6 mm. deep, the campanulate tube 2.1-2.3 mm. long, 1.7-2 mm. in diameter, the subulate teeth 0.8-1.5 mm. long, the whole becoming papery, marcescent unruptured; petals ochroleucous, the banner veined with dull brownish-lilac; banner abruptly recurved through 90° or sometimes further, broadly obovate-cuneate, 5.3-6.8 mm. long, 4-5.4 mm. wide; wings (0.7 mm. longer to 0.7 mm. shorter than the banner) 5.7-6.8 mm. long, the claws 2-2.5 mm., the oblong-obovate, obtuse blades 4-4.4 mm. long, 1.9-2.9 mm. wide, both incurved but the left one further and more abruptly than the right; keel 4-5 mm. long, the claws 1.8-2.3 mm., the halfcircular blades 2.4—3 mm. long, 1.7-1.9 mm. wide, abruptly incurved through 110-130° to the deltoid, often obscurely porrect apex; anthers 0.35-0.5 mm. long; pod pendulous, stipitate, the stipe (1.5) 2—4 mm. long, the body lance- or oblong- ellipsoid, usually a trifle incurved, 7-12 mm. long, (2) 2.5-3.5 (3.8) mm. in diameter, cuneate at both ends or truncately contracted at base into the stipe, cuspidate at apex, triquetrously compressed with nearly flat lateral and openly sulcate dorsal faces, keeled ventrally by the prominent suture, the thin, green but rather densely strigulose valves becoming papery, stramineous, delicately cross- reticulate, inflexed as a not quite complete septum 0.7-1.2 mm. wide; dehiscence apical and through the ventral suture; ovules 6-12; seeds brown or greenish-brown, sometimes purple-speckled, smooth but dull, 1.5-1.8 mm. long.—Collections: 4 (i); representative: Nelson & Macbride 2987 (CAS, GH, MO, NY); R. J. Davis 3088 (IDS).

    Sandy flats and hollows among boulders, sometimes with sagebrush, on basalt, 4750-4900 feet, known only from a restricted area in the foothills of the Sawtooth Range, around Carey and Picabo, Blaine County, Idaho.—Map No. 52.—May to July.

    Astragalus oniciformis (resembling A. Mulfordae, referred by Rydberg to the genus Onix) Barneby in Leafl. West. Bot. 8: 122. 1957.—"Idaho: ... Blaine County... elevation 4750 ft., east of Picabo, 21 June 1947, fl. & fr., Ripley & Barneby 8795 ..."--Holotypus, CAS! isotypi, IDS, NY, RM, RSA, UTC, WTU!

    The Picabo milk-vetch, A. oniciformis, is a puzzling species, introduced at this point on account of its technical similarity to A. misellus, although in many important features of its anatomy and deportment more suggestive of A. Mulfordae of the following section. The shortness of the peduncle in relation to the elongate, distantly flowered raceme-axis, and the tiny flowers with unequally incurved wings are particularly suggestive of sect. Neonix, although the free stipules are at variance with the circumscription of that group. The likeness to A. Mulfordae is immediately striking, but on close observation A. oniciformis is found to differ in several fine details other than the stipules, as brought out in the following key.

    1. Vesture of sinuous or incurved hairs; petals ochroleucous, the banner veined with brownish-purple; leaflets oblong-elliptic, all jointed; body of the pod 6-12 mm. long; ovules 6-12; seeds 1.5—1.8 mm. long; Blaine County, Idaho, at ± 4800 ft. elevation          A. oniciformis

    1. Vesture of appressed, straight hairs; petals white; leaflets mostly linear, the terminal one of some upper leaves nearly always continuous with the rachis; body of the pod (9)10-16 mm. long; ovules (11)12-16; seeds 2-2 6 mm long; Washington and Ada Counties, Idaho, and adjoining Oregon, at 2200-2800 ft. elevation                                             A. mulfordae