Astragalus Castetteri

  • Title

    Astragalus Castetteri

  • Authors

    Rupert C. Barneby

  • Scientific Name

    Astragalus castetteri Barneby

  • Description

    48. Astragalus Castetteri

    Caulescent perennial with a woody taproot and shallowly buried root-crown or caudex, strigulose nearly throughout with fine, straight, appressed hairs up to 0.3-0.45 mm. long, the herbage green or cinereous when young, the leaflets more or less bicolored, brighter green but either glabrous or thinly pubescent above; stems several or numerous, diffuse and incurved-ascending, 2—4 dm. long, leafless and subterranean for a space of 1—12 cm., spurred or shortly branched at 1—5 nodes preceding the first peduncle, floriferous upward from near or a little below the middle; stipules 2—7 mm. long, strongly dimorphic, the lowest ones amplexicaul and connate into a papery, pallid or brownish, subtruncate or obtusely bidentate, eventually glabrous and fragile sheath, the upper ones longer, herbaceous, deltoid-acuminate or lanceolate, amplexicaul but only very shortly or obscurely connate, strigulose dorsally, the blades mostly erect; leaves (3.5) 5—10 cm. long, petioled but the uppermost only shortly so, with (11) 13—25 oblong-elliptic or (in some lower leaves) obovate, emarginate, flat or loosely folded leaflets (2.5) 5-14 mm. long; peduncles rather stout, 4.5—8 cm. long, a little longer or shorter than the leaf; racemes 8—18-flowered, rather dense at early anthesis, the flowers early spreading and declined in age, the axis not greatly elongating, 2—5 cm. long in fruit; bracts submembranous, narrowly lance-acuminate, 2—4 mm. long; pedicels at anthesis slender, ascending, 0.8—1.5 mm. long, in fruit a little thickened, recurved, or straight and divaricate, 2-3.5 mm. long, tardily disjointing with the fruit; bracteoles 0; calyx 7.9—10 mm. long, loosely strigulose or villosulous with incurved and ascending, black or mixed black and white hairs, the strongly oblique disc 1.2-1.6 mm. deep, the deeply campanulate or subcylindric, subtumid, purple- tinged tube 5.8-7.2 mm. long, 3.5-4 mm. in diameter, the subulate teeth 2-3.5 mm. long, the whole becoming papery, marcescent; petals pale purple, drying dull lavender; banner recurved through 45°, broadly rhombic-oblanceolate or -spatulate, openly notched, 14-18.7 mm. long, 7.4-9 mm. wide; wings 14.2-17.6 mm. long, the claws 7-8.1 mm., the narrowly oblong-elliptic, obtuse, almost straight blades 8-10.2 mm. long, 2.4-2.8 mm. wide; keel 11-14.4 mm. long, the claws 6.2-7.8 mm., the half-obovate blades 5.2-7.6 mm. long, 2.8-3.1 mm. wide, abruptly incurved through 90° to the exactly or bluntly deltoid apex; anthers 0.650.75 mm. long; pod widely spreading or declined, stipitate, the stipe 1.5—3 mm. long set at an oblique angle to the subsymmetrically ovoid-acuminate or -ellipsoid, bladdery-inflated body, the body 2.2-2.8 cm. long, 1.1-1.6 cm. in diameter, cuneately contracted at base into the stipe, narrowed distally into a triangular-acuminate, laterally compressed, almost erect beak 4-8 mm. long, otherwise subterete but shallowly sulcate ventrally, the sutures both filiform, the thin, red-mottled, loosely strigulose or subvillosulous valves becoming papery, stramineous, subdiaphanous, somewhat lustrous, finely cross-reticulate, smooth within, not inflexed, the funicular flange 0.7 mm. wide; dehiscence apical, through the beak; ovules 22-26; seeds dark brown, minutely pitted or almost smooth, dull, 2.2-2.8 mm. long.— Collections: 3 (o); representative: Hershey 2007 (RM).

    Dry slopes in piñon forest, 5000-5500 feet, locally common but known only from the west slope of the San Andres Mountains (Ash Canyon; Rope Springs), northeastern Dona Ana County, New Mexico.—Map No. 18.—April and May.

    Astragalus Castetteri (Edward Franklin Castetter, 1891- ), sp. nov., A. Hallii et praesertim ejus var. fallaci (Wats.) Barneby arete affinis, a var. fallacis formis legumine vesicario praeditis differt dentibus calycinis elongatis (2-3.5 nec 0.7-2 mm. longis), legumine ipso maxime inflato (1.1-1.6 cm. nec 0.7-1.2 cm. diametro) striguloso nec villosulo, insuper anthesi vernali (Aprili Maioque mensibus) nec solstitial! et serotino (in Septembrem usque). —New Mexico: common under pinon trees, Rope Springs, Dona Ana County, April 27, 1937, fl. & fr. jun., A. L. Hershey 2005.—Holotypus, RM! isotypi 6 distributuri. Ash Canyon, San Andres Mountains, Dona Ana County, May 24, 1952, fr. maturo, E. F. Castetter 5964. Cotypus in herb. Univ. New Mex!

    The Castetter milk-vetch is closely related to the typical phase of the variable A. Hallii var. fallax, which it resembles in the thin texture and inflation of the pod. It may have arisen from var. fallax through a mutation which involved some small morphological modifications coinciding with a reversion to vernal flowering. The species first became known to me through Professor Castetter’s specimen cited above as the fruiting cotypus, but it was collected first by Hershey in 1937. I am indebted to Professor Porter for permission to analyze the unmounted type-set in the Rocky Mountain herbarium that he had set aside for further study.