Cinchona scrobiculata Humb. & Bonpl.
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Authority
Andersson, Lennart. 1998. A revision of the genus
(Rubiaceae--Cinchoneae). Mem. New York Bot. Gard. 80: 1-75. -
Family
Rubiaceae
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Scientific Name
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Description
Species Description - Tree, to 12 m tall. Young shoots glabrous or sparsely hirsute. Stipules 1.8-2.8 X ca. 0.7 cm, ± tongue-shaped, sparsely hirtellous; petioles 0.4-2 cm long, glabrous; leaf blades papyraceous to (usually) chartaceous when dry, 13-25 X 7.2-11 cm, elliptic or ovate to slightly obovate (length:width ratio 1.5-2.4), round to acute at apex, sometimes slightly acuminate, rounded to cuneate or attenuate at base, secondary veins 11-14 pairs, ± prominulous above, prominent beneath, tertiary veins level on both sides, ± obscure above, rather distinct beneath, adaxial surface matte to (usually) ± glossy, glabrous, abaxial surface usually glabrous throughout (except for domatia), or sparsely puberulous on midrib, or rarely coarsely hirsute throughout, most densely so on midrib and veins, hairs ca. 1 mm long. ± distinct tuft domatia present, especially in vein axils of the proximal part of the blade. Calyx 2-2.4 mm with the triangular lobes 0.5-0.8 mm long, outside moderately to rather densely puberulous, inside rather densely puberulous ± throughout, without colleters; corolla tube ca. 8 mm, villous inside above stamen attachments, lobes ca. 2.9 mm; stamens attached 2.8 mm (ca. 35% of tube length) above base of corolla tube in short-styled flowers (long-styled ones seen in bud only), filaments ca. 4 mm in short-styled flowers, ca. 1.3 mm in long-styled ones, anthers ca. 2.4 X 0.5 mm; style ca. 2.5 mm in short-styled flowers, >6.7 mm in long-styled ones, stigma branches ca. 1.7 mm, ovary 1.6-1.9 X 1.6-2.1 mm, densely villosulous outside, disk glabrous. Capsules 5-18 X 3-6 mm, acropetally dehiscent, with chartaceous endocarp ca. 0.15 mm thick. Seeds irregularly elliptic, 3.7-6.3 X 1.5-2 mm, wing margin ± fimbriate, at least in basiscopic part.
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Discussion
Type. Peru. Cajamarca: “Jaen de Bracamoros,” n.d., Bonpland 3614 (lectotype, P, designated here; isolectotypes, B-W 4032, F, G, P). Cinchona umbellulifera Pav. in Howard, I11. Nueva Quinol. Pav. t. [22]. 1862. Type. Peru. Cajamarca: “ex Jaén,” 1804, [Tafalla in] Pavón 584/590 (lectotype, MA, designated here [see Nomenclatural Notes]; supposed isolectotype, G, with number 584). Nomenclatural notes. The specimen designated as lectotype of Cinchona scrobiculata is one out of two at P with an original label in Bonpland’s hand: “3614 / Cascarilla fina / Cinchona / Jaen”; although the scientific name does not occur on these labels, the identity as type material is proved by the vernacular one. The specimen designated as lectotype of C. umbellulifera has four labels attached to the sheet, apparently two pairs of duplicates, one pair numbered 590 and the other 584. There are two branches mounted on the sheet, one fruiting and one sterile; the fruiting one is designated lectotype. The material at G is unambiguously labeled 584. The number 590 occurs also on the lectotype of C. conglomerata (a synonym of C. villosa). One supposed isotype of C. conglomerata is annotated (in a hand different from Pavón’s) with the numbers 590 and 589, the latter probably a misreading for 584, since the figures 4 and 9 are quite similar in Pavón’s hand. The type collections of C. conglomerata and C. umbellulifera obviously became mixed up at some time, presumably prior to distribution by Pavón. Cinchona scrobiculata is characterized by having ± chartaceous leaf blades with tuft domatia, small flowers with the calyx ± densely puberulous throughout inside, and corolla tube villous inside. Cinchona scrobiculata was synonymized under C. pubescens by Standley (1931a, 1936), who had seen the type material at G. Standley (1936) provisionally recognized C. umbellulifera, but said that he had not seen the type.
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Distribution
Distribution and habitat. Known only from the area around Jaén in the Cajamarca department, N Peru (Fig. 14B), and only from the type collections cited above. No details are known about the habitat.
Peru South America|