Castilleja martinii var. clokeyi (Pennell) N.H.Holmgren

  • Authority

    Holmgren, Noel H. 1971. A taxonomic revision of the Castilleja viscidula group. Mem. New York Bot. Gard. 21: 1-63.

  • Family

    Scrophulariaceae

  • Scientific Name

    Castilleja martinii var. clokeyi (Pennell) N.H.Holmgren

  • Description

    Species Description - Stems 1.5-4 (-5) dm high, mostly unbranched, wholly herbaceous above the thick, woody caudex. Leaves 1-4 (-5.5) cm long, the main leaves relatively broad, lanceolate to ovate, rounded or obtuse, rarely acute. Inflorescence relatively broad. Bracts villose-ciliate on the proximal margin. Calyx 16-21 (-28) mm long, the abaxial cleft (5-)6-9 mm deep, the adaxial cleft (6-)7-10(-11) mm deep, the primary lobes 2/5 to 1/2 the calyx length, the segments (1-)2-(-5) mm long, lanceolate to triangular, acute to obtuse, the tube glandular- pubescent and villosulose. Corolla 25-32(-40) mm long; galea more or less as long as the tube, 11-18 (-20) mm long, usually dorsally green, rarely red all over; tube (11-) 13-20(-22) mm long. Stamens often exserted beyond the galea tip. Capsules relatively long, 11-14(-17) mm long. Chromosome number: 2n=48.

  • Discussion

    Castilleja clokeyi Pennell, Proc. Acad. Sci. Philadelphia 89: 420. 1937.

    Type. Nevada, Clark County, Charleston Mountains, Kyle Canyon, gravelly loam, N slope with Pinus scopulorum (Engelm.) Lemmon ( = P ponderosa Laws.) and Populus aurea Tidestr. ( = P tremuloides Michx.), alt. 2425 m (8000 ft), 8 Jul 1936, Clokey d Clokey 7322 (PH 757618, isotypes, BRY, CAS, DS, MO, NY, ORE, OSC, RSA, SD, UC, US, UTC, WTU).

    Since the publication of Pennell's description of Castilleja clokeyi, it has been considered a distinct species endemic to the Charleston ]Mountains. Collections from the Sheep and Panamint Ranges and the Inyo Mountains have heretofore passed for C. applegatei. Until now it has never been compared with the California species C. martinii, which it so closely resembles that it must be regarded as a variety.

    In comparison with C. applegatei var fragilis, C. martinii var clokeyi has a shorter calyx with shorter primary lobes and segments, and has proportionately broader and often rounded leaves. Pennell had written an unpublished varietal epithet on some of the collections from the Panamint Range and the Inyo Mountains, which indicates that he must have realized that the specimens were not exactly the C. applegatei as then accepted.

    There are only two other species of CastiUeja that cross the range of var clokeyi. These are C. chromosa A. Nels., which is of lower elevations and flowers much earlier, and C. linariaefolia Benth., which comes into contact with var clokeyi only at the upper extremes of its altitudinal range. I have observed no evidence from field observations and herbarium specimens of hybridization between these species.

  • Distribution

    Habitat and distribution. Collected mostly from late May to early August, but sometimes as late as September at higher altitudes, from 7000 to 10,200 feet elevation; on dry gravelly slopes, in juniper, pinyon, and cercocaipus woodlands of the foothills, talus, dry meadows, and in open places of ponderosa, bristlecone pine, and aspen forests from middle elevations to timberline. In the Basin Ranges of southern Nevada, from the Quinn Canyon Range, south to the Sheep Range and Charleston Mountains

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