Diplazium lonchophyllum Kunze
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Authority
Mickel, John T. & Smith, Alan R. 2004. The pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1-1054.
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Family
Athyriaceae
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Scientific Name
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Description
Species Description - Rhizomes erect; rhizome scales 4–6 x 1–1.5 mm, black, sclerotic, linear-lanceolate, entire; fronds (15–)45–100 cm; stipes green to stramineous, dark at bases, (3–)14–22(–40) cm x 2–3 mm, 1/4–2/5 the frond length, bases occasionally with black scales; blades chartaceous, mostly pinnate-pinnatifid, 2-pinnate only at bases, (8–)25–50 x (4–)12–20 cm, deltate to ovate-lanceolate, apices pinnatifid, buds absent; rachises glabrous; pinnae stalked (1–)3–8 mm, inequilateral, (8–)15–20 pairs per blade, (2–)8–16 x (1–)2–4(–6) cm, cut 4/5 to the costae or more, bases cuneate to subtruncate, basiscopic segments shorter and more oblique than acroscopic ones, an acroscopic lobe free or nearly so on proximal 1–4 pinna pairs, apices long-acuminate, lobes usually ascending, narrowly triangular to deltate, acute and serrulate at tips; veins free, pinnate, 6–9 pairs per segment, mostly 1-forked; indument adaxially absent or of sparse, papillae less than 0.1 mm long in rachial and costal grooves, blades abaxially glabrous except for brownish hair-like scales 1–2 mm long on costae; sori pinnate in segments, 6–10 pairs per segment (sometimes on both veins of a forking veinlet), with tan to brownish± entire to erose indusia 4–6 x 0.5–1 mm; 2n=82 (Ver), 164 (Chis, Oax).
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Discussion
Type. Mexico. Veracruz: Jalapa, Schiede s.n. (isotype BM, photos MO!, NY!, UC!, US!).
Diplazium inaequilaterum Liebm., Mexic. Bregn. 255 (reprint 103). 1849. Syntypes. Mexico. Veracruz: all collected by Liebmann s.n., “Mirador,” 1/43 [Pl.Mex. 2379, Fl. Mex. s.n.], “circa Mirador,” 3/42 [Pl. Mex. 2379, Fl. Mex. 355], “Mirador,” 7-11/41 [Pl. Mex. 2379, Fl. Mex. s.n.], “Baranca de Jovo,” 5/41 [Pl. Mex. 2380, Fl. Mex. 354, s.n.] (C, photos NY!, UC!, US!, frag. NY!; isosyntypes K!-2 sheets). Diplazium acutale Fe´e, Me´m. Foug. 5: 215. 1852. Type. Mexico. [Veracruz: Mirador] Galeotti 6289 (P!; isotypes BR!, K! RB).
Unverified, Doubtful, or Mistaken Reports. SLP (reported by Smith, 1981, and Mickel & Beitel, 1988, but not verified). Diplazium lonchophyllum is quite variable and, like D. cristatum and D. drepanolobium (q.v.), in need of monographicstudy. It differs from those two species by the proximal pinnae each having a distinct, free or nearly free, basal acroscopic pinnule. Diplazium drepanolobium has longer and wider pinnae that are strongly oblique and falcate, longer rhizome scales, and narrower, subentire indusia. Intermediate specimens (Mickel 1619, 6184, NY) have malformed spores, and Mickel 1619 is tetraploid, with 41II and 75I; Mickel et al., 1966); these represent putative hybrids involving D. lonchophyllum as one parent. In Chiapas, Breedlove 21624A, 22293, 22484, and 27480 (DS) are all more or less intermediate between D. drepanolobium and D. lonchophyllum. Breedlove 33658 (DS), cited as D. prominulum Maxon by Smith (1981) from Chiapas, is D. lonchophyllum. Diplazium lonchophyllum was reported from Nuevo Leo´n by Aguirre-Clavera´n and Arregui´n-Sa´nchez (1988), and a possible specimen is Sa´nchez 603, MEXU. This specimen varies in the direction of D. franconis.
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Distribution
Terrestrial on slopes and along streams; 750- 1550(-2450) m. Mexico; Guat, Hond, Salv, Nic, CR, Pan; Ec. Adams (in Davidse et al., 1995) also cited this species from Colombia.
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