Taxon Details: Tococa croatii Almeda
Taxon Profile:
 | The Plant List |
 | International Plant Name Index |
 | Tropicos |
 | Catalogue of Life |
 | Global Biodiversity Information Facility |
 | JSTOR Types |
| JSTOR |
 | BHL |
 | Encyclopedia of Life |
 | WikiSpecies |
 | Google Scholar |
 | PubMed |
 | Morphbank |
 | IUCN |
 | National Center for Biotechnology Information |
 | Barcode of Life |
Narratives:
Family:
Melastomataceae (Magnoliophyta)
Melastomataceae (Magnoliophyta)
Scientific Name:
Tococa croatii Almeda
Tococa croatii Almeda
Primary Citation:
Proc. Calif. Acad. Sci. ser. 4, 46(5): 148. 1989
Proc. Calif. Acad. Sci. ser. 4, 46(5): 148. 1989
Accepted Name:
This name is currently accepted.
This name is currently accepted.
Description:
Description Author and Date: Fabian A. Michelangeli, January 2010, based on Michelangeli, F. A. (2005). Tococa (Melastomataceae). Flora Neotropica Monographs 98: 1-114.
Type: Panama. Darién: Serranía de Pirre, 1520-1560 m, 27 JUL 1976, Croat 37873 (holotype CAS; Isotype MO).
Description: Shrub, to 0.5-1.5 m tall. Stems with straight, eglandular hairs, the trichomes 1-2 mm long, caducous, the nodes glabrous. Petioles 2.5-12.5 cm long, densely to sparsely pubescent, the trichomes often caducous. Leaves isophyllous or slightly anisophyllous, broadly ovate to cordate, 12-19.5 x 8.0-15.5 cm, apex acuminate, base cordate, to broadly rounded, adaxial surface sparsely to moderately setulose, the trichomes adpressed 0.5-2.0 mm long, abaxial surface moderately to densely glandular pubescent on the primary and secondary veins, the trichomes late caducous to persistent, the surface between the veins fine-setulose, light green, with 3(-4) pairs of secondary veins, basally nervate, membranaceous, crenulate, ciliate; domatia absent. Inflorescence a terminal panicle, of 6-20 (or more) flowers, 10-18 cm long, the axis green, terete. Flowers on pedicels 1.0-4.0 mm long, not flared at the apex, subtended by late caducous bracts, subulate to setiform, 1-1.2 x 0.1-0.2 mm; hypanthium campanulate, 2.0-2.5 mm long, with a mix of sparse glandular trichomes and sessile glands; sepals broadly deltoid, hyaline, 2.0 mm long, calyx teeth mostly fused to the sepals, the subulate apex projecting beyond sepals, 0.25-0.5 mm long; the ring inside the torus with minute glandular trichomes. Petals 5, obovate, 5.0-6.0 x 3.5-5.0 mm, base truncate to obtuse, entire, smooth, glabrous, pink, white, or even bluish. Stamens all of the same size, laterally coherent (connivent) forming a ring around the style, the filaments 2.5-3.5 mm long, glabrous; anthers yellow; the connective without a dorsal-basal blunt tooth, but the base prolonged into a retrorse triangular appendage1.0-1.5 x 1.0 mm; thecae 1.5-2.0 mm long, opening by an upright terminal pore. Ovary 5-locular, terete, 1/2-2/3 inferior, the superior portion spherical, with a corona of short (up to 1.5 mm) glandular setae; style glabrous, 5.5-8.5 mm long; stigma truncate to distally papillate. Fruits globose, 3.5-4.0 mm long, black to deep blue, sparsely glandular setose to glabrous. The seeds not evident through the pericarp, truncate-obovate, 0.5-0.6 mm long, without capitate trichomes in the raphal area, with sculpturing, anticlinal walls s-shaped, periclinal walls flat to slightly convex, boundaries between periclinal walls flat. Chromosome number n= unknown.
Habitat and Distribution: Southestern Panama. Growing in the understory of Tropical Montane forests (from 1300 to 1700 m) in the Darién, near the border with Colombia, where it might also occur.
Taxonomy and Systematics: For details on the systematic position of T. croatii see discussion under T. symphyandra.
Description Author and Date: Fabian A. Michelangeli, January 2010, based on Michelangeli, F. A. (2005). Tococa (Melastomataceae). Flora Neotropica Monographs 98: 1-114.
Type: Panama. Darién: Serranía de Pirre, 1520-1560 m, 27 JUL 1976, Croat 37873 (holotype CAS; Isotype MO).
Description: Shrub, to 0.5-1.5 m tall. Stems with straight, eglandular hairs, the trichomes 1-2 mm long, caducous, the nodes glabrous. Petioles 2.5-12.5 cm long, densely to sparsely pubescent, the trichomes often caducous. Leaves isophyllous or slightly anisophyllous, broadly ovate to cordate, 12-19.5 x 8.0-15.5 cm, apex acuminate, base cordate, to broadly rounded, adaxial surface sparsely to moderately setulose, the trichomes adpressed 0.5-2.0 mm long, abaxial surface moderately to densely glandular pubescent on the primary and secondary veins, the trichomes late caducous to persistent, the surface between the veins fine-setulose, light green, with 3(-4) pairs of secondary veins, basally nervate, membranaceous, crenulate, ciliate; domatia absent. Inflorescence a terminal panicle, of 6-20 (or more) flowers, 10-18 cm long, the axis green, terete. Flowers on pedicels 1.0-4.0 mm long, not flared at the apex, subtended by late caducous bracts, subulate to setiform, 1-1.2 x 0.1-0.2 mm; hypanthium campanulate, 2.0-2.5 mm long, with a mix of sparse glandular trichomes and sessile glands; sepals broadly deltoid, hyaline, 2.0 mm long, calyx teeth mostly fused to the sepals, the subulate apex projecting beyond sepals, 0.25-0.5 mm long; the ring inside the torus with minute glandular trichomes. Petals 5, obovate, 5.0-6.0 x 3.5-5.0 mm, base truncate to obtuse, entire, smooth, glabrous, pink, white, or even bluish. Stamens all of the same size, laterally coherent (connivent) forming a ring around the style, the filaments 2.5-3.5 mm long, glabrous; anthers yellow; the connective without a dorsal-basal blunt tooth, but the base prolonged into a retrorse triangular appendage1.0-1.5 x 1.0 mm; thecae 1.5-2.0 mm long, opening by an upright terminal pore. Ovary 5-locular, terete, 1/2-2/3 inferior, the superior portion spherical, with a corona of short (up to 1.5 mm) glandular setae; style glabrous, 5.5-8.5 mm long; stigma truncate to distally papillate. Fruits globose, 3.5-4.0 mm long, black to deep blue, sparsely glandular setose to glabrous. The seeds not evident through the pericarp, truncate-obovate, 0.5-0.6 mm long, without capitate trichomes in the raphal area, with sculpturing, anticlinal walls s-shaped, periclinal walls flat to slightly convex, boundaries between periclinal walls flat. Chromosome number n= unknown.
Habitat and Distribution: Southestern Panama. Growing in the understory of Tropical Montane forests (from 1300 to 1700 m) in the Darién, near the border with Colombia, where it might also occur.
Taxonomy and Systematics: For details on the systematic position of T. croatii see discussion under T. symphyandra.
Flora and Monograph Treatment(s):
Tococa croatii Almeda: [Article] Michelangeli, Fabián A. 2005. (Melastomataceae). Fl. Neotrop. Monogr. 98: 1-114.
Tococa croatii Almeda: [Article] Michelangeli, Fabián A. 2005.