Taxon Details: Tococa racemifera Wurdack
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Narratives:
Family:
Melastomataceae (Magnoliophyta)
Melastomataceae (Magnoliophyta)
Scientific Name:
Tococa racemifera Wurdack
Tococa racemifera Wurdack
Primary Citation:
Phytologia 48: 247. 1981
Phytologia 48: 247. 1981
Accepted Name:
This name is currently accepted.
This name is currently accepted.
Description:
Description Author and Date: Fabian A. Michelangeli, January 2010, based on Michelangeli, F. A. (2005). Tococa (Melastomataceae). Flora Neotropica Monographs 98: 1-114.
Type: Colombia. Antioquia: wet forest, 3 km from Planta Providencia, 28 Km SW of Zaragoza, Río Anori Valley, 400-700 m, 21 FEB 1977, Alverson, White & Shepherd 66 (holotype US; isotypes NY [2], WIS[n.v.]).
Description: Shrub, up to 2-3 m tall. Stems with a mix of sparsely non glandular trichomes 2.0-4.0 mm long glandular trichomes up to 1.5 mm long and sparse stallate hairs, persistent, the nodes glabrous. Leaves strongly anysphyllous, Petioles 0.9-1.3 cm long, pubescence as in the stems. Leaves anisophyllous, adaxial and abaxial surfaces sparsely setose, the trichomes up to 1. 5 mm long, membranaceous, light green, the borders crenulate to serrulate; the large leaf of each pair with 2 pairs of secondary veins, basally nervate, elliptic, 17-30 (-40) x 7.5-14 (-16) cm, apex acuminate, the terminal apex 1.2-2.0 cm long, base acute to obtuse; domatia present, free or or with the apex slightly immersed in the leaf blade, 1.4-1.7 x 0.7-0.9 cm; the smaller leaf of each pair with petioles 0.2-0.3 cm long, without domatia, or if present poorly developed, ovate to elliptic, the base round to cordate, with 1(-2) pairs of secondary veins, basally nervate. Inflorescence a slender raceme, terminal becoming axillary by overgrowth of the axillary meristem, 9-26 cm long, with numerous flowers, the axis terete, green, densely pubescent with the trichomes redish; flower pedicels sessile; bracts persistent, hypanthium conical, slightly 10-costate, 2.6-2.8 mm long, densely glandulose-setose on the ridges, the trichomes up to 1 mm long, densely stellate in between the ridges; sepals partly fused, the terminal lobes deltoid, 0.3-0.5 mm long, calyx teeth acute, with 2-3 apical setae up to 1.5 mm long, 0.7-1.0 mm long;; the ring inside the torus glabrous. Petals 5, emarginate, 2.9-3.1 x 0.8-0.9 mm, smooth, glabrous, white. Stamens all of the same size, the filaments 3.0-3.1 mm long, glabrous; anthers yellow; the connective with a dorsal-basal blunt tooth; thecae 2.8-3.1 mm long, opening by a dorsally inclined small pore. Ovary 3-4-locular, 2/3 inferior, the superior portion sulcate, the apex glabrous; style glabrous, 8.0-9.0 mm long; stigma capitate, 0.5-0.6 mm wide. Fruits globose, 3.5-4.5 mm long, black, sparsely glandular pubescent. The seeds evident through the pericarp, truncate-obovate, 0.4 mm long, without capitate trichomes in the raphal area, with sculpturing, anticlinal walls s-shaped, periclinal walls flat, boundaries between periclinal walls flat. Chromosome number n= unknown.
Habitat and Distribution: Endemic to the deparment of Antioquia, Colombia. Growing in forests up to 1000 m.
Taxonomy and Systematics: Tococa racemifera is known from only three colletions from two localities in Antioquia, Colombia. The morphology of the inflorescences makes it a very distinct species. However, given some characters of its floral morphology, and mostly due to the anatomy of the cells of the seed testa, its affinities to other species of the Miconieae are not clear. Other species of Tococa have inflorescences that are laterally compressed (T. pachystachya and T. spadiciflora), but differences in the bracteoles and shape of the peduncle appear to negate a common origin for this characater.
Description Author and Date: Fabian A. Michelangeli, January 2010, based on Michelangeli, F. A. (2005). Tococa (Melastomataceae). Flora Neotropica Monographs 98: 1-114.
Type: Colombia. Antioquia: wet forest, 3 km from Planta Providencia, 28 Km SW of Zaragoza, Río Anori Valley, 400-700 m, 21 FEB 1977, Alverson, White & Shepherd 66 (holotype US; isotypes NY [2], WIS[n.v.]).
Description: Shrub, up to 2-3 m tall. Stems with a mix of sparsely non glandular trichomes 2.0-4.0 mm long glandular trichomes up to 1.5 mm long and sparse stallate hairs, persistent, the nodes glabrous. Leaves strongly anysphyllous, Petioles 0.9-1.3 cm long, pubescence as in the stems. Leaves anisophyllous, adaxial and abaxial surfaces sparsely setose, the trichomes up to 1. 5 mm long, membranaceous, light green, the borders crenulate to serrulate; the large leaf of each pair with 2 pairs of secondary veins, basally nervate, elliptic, 17-30 (-40) x 7.5-14 (-16) cm, apex acuminate, the terminal apex 1.2-2.0 cm long, base acute to obtuse; domatia present, free or or with the apex slightly immersed in the leaf blade, 1.4-1.7 x 0.7-0.9 cm; the smaller leaf of each pair with petioles 0.2-0.3 cm long, without domatia, or if present poorly developed, ovate to elliptic, the base round to cordate, with 1(-2) pairs of secondary veins, basally nervate. Inflorescence a slender raceme, terminal becoming axillary by overgrowth of the axillary meristem, 9-26 cm long, with numerous flowers, the axis terete, green, densely pubescent with the trichomes redish; flower pedicels sessile; bracts persistent, hypanthium conical, slightly 10-costate, 2.6-2.8 mm long, densely glandulose-setose on the ridges, the trichomes up to 1 mm long, densely stellate in between the ridges; sepals partly fused, the terminal lobes deltoid, 0.3-0.5 mm long, calyx teeth acute, with 2-3 apical setae up to 1.5 mm long, 0.7-1.0 mm long;; the ring inside the torus glabrous. Petals 5, emarginate, 2.9-3.1 x 0.8-0.9 mm, smooth, glabrous, white. Stamens all of the same size, the filaments 3.0-3.1 mm long, glabrous; anthers yellow; the connective with a dorsal-basal blunt tooth; thecae 2.8-3.1 mm long, opening by a dorsally inclined small pore. Ovary 3-4-locular, 2/3 inferior, the superior portion sulcate, the apex glabrous; style glabrous, 8.0-9.0 mm long; stigma capitate, 0.5-0.6 mm wide. Fruits globose, 3.5-4.5 mm long, black, sparsely glandular pubescent. The seeds evident through the pericarp, truncate-obovate, 0.4 mm long, without capitate trichomes in the raphal area, with sculpturing, anticlinal walls s-shaped, periclinal walls flat, boundaries between periclinal walls flat. Chromosome number n= unknown.
Habitat and Distribution: Endemic to the deparment of Antioquia, Colombia. Growing in forests up to 1000 m.
Taxonomy and Systematics: Tococa racemifera is known from only three colletions from two localities in Antioquia, Colombia. The morphology of the inflorescences makes it a very distinct species. However, given some characters of its floral morphology, and mostly due to the anatomy of the cells of the seed testa, its affinities to other species of the Miconieae are not clear. Other species of Tococa have inflorescences that are laterally compressed (T. pachystachya and T. spadiciflora), but differences in the bracteoles and shape of the peduncle appear to negate a common origin for this characater.
Flora and Monograph Treatment(s):
Tococa racemifera Wurdack: [Article] Michelangeli, Fabián A. 2005. (Melastomataceae). Fl. Neotrop. Monogr. 98: 1-114.
Tococa racemifera Wurdack: [Article] Michelangeli, Fabián A. 2005.
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