Taxon Details: Megalastrum longiglandulosum R.C.Moran & J.Prado
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Family:

Dryopteridaceae (Pteridophyta)
Scientific Name:

Megalastrum longiglandulosum R.C.Moran & J.Prado
Accepted Name:

This name is currently accepted.
Description:

by: R.C. Moran, J. Prado

Type: Costa Rica. Puntarenas: Osa Peninsula, ca. 4 mi W of Rincón de Osa, near airfield, 8°42'N, 83°31'W, 30 m, 4-7 Jun 1968, W. C. Burger & R. G. Stolze 5421 (Holotype: F-2 sheets; isotypes: CR, GH-2 sheets, US).

Description: Rhizomes compact, ascesnding (from original description); leaves up to 1.5 m long; scales of the petiole bases 15-25 × 0.3-0.6 mm, linear, spreading-ascending, brown, shiny, denticulate on the margins; laminae up to 0.9 m long, 3-pinnate-pinnatisect at base, 2-pinnate-pinnatisect medially; basal pinnae 15-40 cm long, strongly inequilateral (elongated basiscopically); pinna rachises abaxially glandular (both stipitate and sessile glands present), densely pubescent, scaly, the scales 0.8-1.0 × ca. 0.2 mm, linear-lanceolate, non-bullate, mostly appressed, shiny, brown, sparsely denticulate, adaxially densely pilose, sparsely glandular, sparsely scaly, hairs ca. 1.0 mm long, 6-8-celled, spreading, glands spherical and sessile and stalked, ca. 0.2 mm long; costules sparsely pilose, glandular, hairs 0.3-0.5 mm long, 3-5-celled, erect, the glands both sessile and spherical, and stalked-glandular, adaxially like the pinna rachises adaxially; laminar tissue between the veins abaxially densely glandular, the glands 0.2-0.3 mm long, 2- or 3-celled, erect, capitate, the terminal cell yellow, shiny, these mixed with hairs 0.2-0.3 mm long, acicular, 1- or 2-celled, adaxially without hairs, non-glandular or with a few sparse sessile spherical glands; veins visible on both surfaces, abaxially with indumenta like the laminar tissue, adaxially sparsely pilose, glandular, the hairs 0.4-0.8 mm long, 4-6-celled, glands sessile, spherical; hydathodes evident; lamina margins pubescent, hairs 0.3-0.5 mm long, 3- or 4-celled, ascending, acicular, non-glandular; indusia absent.

Distribution: Costa Rica, endemic to the Osa Peninsula; wet forests, 30-600 m.

Comments: Megalastrum longiglandulosum bears dense, erect, long-stalked capitate-glandular hairs between the veins abaxially (the specific epithet refers to this character). In this it resembles M. gilbertii (Clute) R. C. Moran, J. Prado & Labiak, a West Indian species, but it differs by filiform scales and densely pubescent pinna rachises abaxially (M. gilbertii has wider scales and hairless pinna rachises abaxially). The size of the glandular hairs on the laminar tissue between the veins abaxially is also different: those on M. glandulosum are up to 0.3 mm long, whereas those on M. gilbertii are ca. 0.1 mm long. Two Central American species, Megalastrum lunense and M. atrogriseum, also have long-stalked capitate-glandular hairs on the laminae abaxially, but their hairs are usually mixed with many non-glandular, acicular hairs of variable length. These two species also differ from M. longiglandulosum by the scales on the abaxial surfaces of the pinna rachises: the scales of M. lunense are ovate-lanceolate and flaccid, and those of M. atrogriseum are linear, firm (not twisted or tortuous), and golden brown.