Monographs Details:
Authority:
Pedraza-Peñalosa, Paola & Luteyn, James L. 2010. New species of Ericaceae (Vaccinieae) from the Andes of Bolivia and Peru. Brittonia. 62 (1): 39-56.
Pedraza-Peñalosa, Paola & Luteyn, James L. 2010. New species of Ericaceae (Vaccinieae) from the Andes of Bolivia and Peru. Brittonia. 62 (1): 39-56.
Family:
Ericaceae
Ericaceae
Description:
Description - Epiphytic shrubs. Twigs terete to subterete, often bluntly angles, puberulous, the hairs simple, some unicellular and eglandualr, or multicellular, glandular and deciduous; mature stems terete, the bark fissured, brown, glabrescent, the hairs simple, unicellular and eglandular; prophylls 2, opposite, ovate, obscure, < 0.5 mm long, the apex obstuse. Leaves alterante, spirally arranged; petioles terete, 0.8-1.5 mm long, glaberscent with simple deciduous hairs, both glandular and muticellular long, and, eglandular and unicellular; blades subcoriaceous, elliptic to ovate- elliptic, 1.2 - 2.5 x (0.6-) 0.7 -1.1 cm, the base cuneate to obtuse, to bluntly acute, glabrescent above with simple hairs, some glandular and mutlicellular, others eglandular and unicellular, glabrescent beneath with scattered simple, multicellular and glandular hairs; venation obscurely 3-plinerved, the midrib conspicuous and slightly impressed above, raised beneath, the secondary and tertiary nerves obscure above, slightly raised beneath. Axillary solitary flowers; bracts and floral bract 3-5, undifferen- tiated, ovate to reniform, 0.4-0.5 mm long, the margin eciliate, the apex obtuse, glabrous on both surfaces; pedicels inconspicuously articu- late with calyx, 1.3-2.5 mm long, glabrescent with scattered simple, multicellular and glan- dular hairs; bracteoles 2, distinct, persistent, inserted towards base of pedicel, subopposite, transversely elliptic, 0.6-1x1.2-1.8 mm, the margin eciliate, the apex truncate, glabrous on both surfaces. Flowers 4-merous, diplostemonous. Calyx campanulate overall, conspicuously winged, 3.2-3.7 mm long, glabrescent over the lobes with hairs simple, multicellular and glandular, and also simple, unicellular and eglandular, the aestivation unknown; tube obconic, 2-2.2 mm long; limb 1.2-1.6 mm long; lobes deltate, 0.6-0.8x1.2 mm, the margin eciliate, the apex acute; sinuses broadly acute. Corolla apparently bistratose, pale pink, cylindric, 8 mm long, 3 mm diam., glabrous without but puberulous within, the hairs simple, unicellular and eglandular, the aestivation apparently valvate; lobes deltate, 1-1.2 x 1.8 - 2mm, the apex acute. Stamens 8, equal, included, 6.8-7 mm long; filaments equal, distince, long triangular and straight, 3.2 - 3.5 mm long, glabrescent in distal half on both surfaces but more abundantly adaxially, the hairs simple, unicellular and eglandular; anthers equal, distinct, 3.8 -4 mm long; thecae 1.3 -1.5 mm long with basal appendage, smooth, cells with polygonal shape; tubules 2, distinct, 2.7 mm long, minutely papillate especially at base, dehiscing by introrse slits, ca. 1.2 mm long. Ovary 4-locular, placentation axile; style included, 7-8 mm long, Berry unknown.
Description - Epiphytic shrubs. Twigs terete to subterete, often bluntly angles, puberulous, the hairs simple, some unicellular and eglandualr, or multicellular, glandular and deciduous; mature stems terete, the bark fissured, brown, glabrescent, the hairs simple, unicellular and eglandular; prophylls 2, opposite, ovate, obscure, < 0.5 mm long, the apex obstuse. Leaves alterante, spirally arranged; petioles terete, 0.8-1.5 mm long, glaberscent with simple deciduous hairs, both glandular and muticellular long, and, eglandular and unicellular; blades subcoriaceous, elliptic to ovate- elliptic, 1.2 - 2.5 x (0.6-) 0.7 -1.1 cm, the base cuneate to obtuse, to bluntly acute, glabrescent above with simple hairs, some glandular and mutlicellular, others eglandular and unicellular, glabrescent beneath with scattered simple, multicellular and glandular hairs; venation obscurely 3-plinerved, the midrib conspicuous and slightly impressed above, raised beneath, the secondary and tertiary nerves obscure above, slightly raised beneath. Axillary solitary flowers; bracts and floral bract 3-5, undifferen- tiated, ovate to reniform, 0.4-0.5 mm long, the margin eciliate, the apex obtuse, glabrous on both surfaces; pedicels inconspicuously articu- late with calyx, 1.3-2.5 mm long, glabrescent with scattered simple, multicellular and glan- dular hairs; bracteoles 2, distinct, persistent, inserted towards base of pedicel, subopposite, transversely elliptic, 0.6-1x1.2-1.8 mm, the margin eciliate, the apex truncate, glabrous on both surfaces. Flowers 4-merous, diplostemonous. Calyx campanulate overall, conspicuously winged, 3.2-3.7 mm long, glabrescent over the lobes with hairs simple, multicellular and glandular, and also simple, unicellular and eglandular, the aestivation unknown; tube obconic, 2-2.2 mm long; limb 1.2-1.6 mm long; lobes deltate, 0.6-0.8x1.2 mm, the margin eciliate, the apex acute; sinuses broadly acute. Corolla apparently bistratose, pale pink, cylindric, 8 mm long, 3 mm diam., glabrous without but puberulous within, the hairs simple, unicellular and eglandular, the aestivation apparently valvate; lobes deltate, 1-1.2 x 1.8 - 2mm, the apex acute. Stamens 8, equal, included, 6.8-7 mm long; filaments equal, distince, long triangular and straight, 3.2 - 3.5 mm long, glabrescent in distal half on both surfaces but more abundantly adaxially, the hairs simple, unicellular and eglandular; anthers equal, distinct, 3.8 -4 mm long; thecae 1.3 -1.5 mm long with basal appendage, smooth, cells with polygonal shape; tubules 2, distinct, 2.7 mm long, minutely papillate especially at base, dehiscing by introrse slits, ca. 1.2 mm long. Ovary 4-locular, placentation axile; style included, 7-8 mm long, Berry unknown.
Discussion:
Phenology. - The type was flowering in July
Etymology - Themistoclesia siranensis is named for the Cerros del Sira, an isolated mountain range in the Department of Huánuco, Peru.Themistoclesia siranensis is characterized by bluntly acute to obtuse leaf apices, solitary flowers, pedicels articulate with the calyx, subsessile, 4-merous flowers, conspicuously winged calyces, and large corollas that are puberulous within. Themistoclesia siranensis resembles T. tunquiniensis as they both have leaves with similar shapes and sizes, 4- merous flowers and subsessile, and corollas puberulous within. The main differences between these species are numerous and are summarized in Table I.