Monographs Details:
Authority:
Maguire, Bassett. 1978. The botany of the Guayana Highland--part X. Mem. New York Bot. Gard. 29: 1-288.
Maguire, Bassett. 1978. The botany of the Guayana Highland--part X. Mem. New York Bot. Gard. 29: 1-288.
Family:
Ericaceae
Ericaceae
Description:
Distribution and Ecology - Distribution. VENEZUELA. Edo. Bolivar: Cerro Auyan-tepui, Alto Caroni, 1600 m alt, Jan 1949, Cardona 2635; Ptari-tepui, vicinity of Cave Rock, 1600-2000 m alt, 14-19 Aug 1970, Moore et al 9734; Chimanta Massif, Central Section, vicinity of Upper Falls of Rio Tirica above Summit Camp, 1940-1950 m alt, 7 Feb 1955, Steyermark & Wurdack 519, 553; Steyermark Falls on upper reaches of Rio Tirica, 1700-1750 m alt, 3 Jul 1953, Steyermark 76012; Apacara-tepui, between Camp 3 and main line of sandstone bluffs, 1210-1600 m alt, 17 Apr 1953, Steyermark 75086; Chimanta-tepui (Torono-tepuí), above valley of Rio Tirica, 1000-1700 m alt, 16 May 1953, Steyermark 75454; Gran Sabana, Ilu-tepui, Camp 1, 1000 m alt, 10 Mar 1952, Maguire 33334; Rio Aponguao drainage ca km 146 south of El Dorado, 1350-1400 m alt, 21 Dec 1970, Steyermark et al 104214; Cerro Venamo, southwest part near Rio Venamo, 950-1150 m alt, 29-30 Dec 1963, Steyermark et al 92404; Cerro Venamo, northwest slopes, 1100-1300 m alt, 14 Apr 1960, Steyermark & Nilsson 134; Cerro Venamo region, between campamento 125 and km 127, wooded ridge of Fila de La Danta, 1200 m alt, 15-17 Apr 1960, Steyermark & Nilsson 207; Meseta del Jaua, Cerro Jaua, southwest portion, along tributary of Rio Marajano, 1750-1800 m alt, 22-28 Feb 1974, Steyermark et al 109659; Cerro Guanacoco, northwest portion, 1450 m alt, 3 Mar 1974, Steyermark et al 109756; Salto Angel, 1200 m alt, 18 Aug 1968, Foldats 7211; slopes of Cerro Uei, 860-1050 m alt, 27-28 Dec 1970, Steyermark et al 104560; Rio Cuyuni drainage, at km 142-143 S of El Dorado, 1300-1380 m alt, 22-28 Dec 1970, Steyermark et al 104260; Sierra W of waterfall of Rio Chixanan, 80 km southeast of El Dorado, 700 m alt, 29 Aug 1961, Steyermark 89608; Sierra Pacaraima, headwaters of Rio Paragua, 1400 m alt, 4-5 Mar 1973, Steyermark 107232; Cerro Guaiquinima, Alto Rio Paragua, 300-500 m alt, Cardona 901, 1077; slopes of Cerro Opacara, 900 m alt, 26 Jun 1946, Cardona 1601. Territorio de Amazonas: Cerro Huachamacari, Rio Cunucunuma, base of East Ridge, 1500 m alt, 8 Dec 1950, Maguire et al 30044; Cerro Marahuaca, 1600 m alt, 10 May 1949, Maguire & Maguire 29174; Cerro de la Neblina, just S of Camp 3, 650-700 m alt, 23 Dec 1953, aguire et al. 36812; Cerro Yapacana, 825 m alt, 4 May 1970, Steyermark & Bunting 103086; Serrania Yutaja, Rio Manapiare, Northwest Ridge, 1400 m alt, 11 Feb 1953, Maguire & Maguire 35160; Cerro Yutaje, left fork of Caño Yutaje, 1250 m alt, 12 Feb 1953, Maguire & Maguire 35201; Serrania Paru, around Laguna Asisa, Cerro Asisa, 1310 m alt, 7 May 1973, Hoyos &. Morillo 54; region of Duida and Marahuaca, Campo Benitez, Apr-Jun 1950, Barnes 55572; Cerro Sipapo, Forest Phelps Camp to North Savanna, 17 Dec 1948, Maguire & Politi 27753; Cerro Duida, Culebra Creek Basin, 1400 m alt, 23 Apr 1949, Maguire & Maguire 29143; Cerro Duida, near escarpment at 1400 m alt, 23 Apr 1949, Maguire & Maguire 29145; Cerro Duida, ridge W of Caño Culebra, 1700 m alt, 22 Nov 1950, Maguire et al 29707; Cerro de la Neblina, Rio Yatua, trail between Camp 2 and 3, 500-700 m alt, 7 Nov 1957, Maguire et al 41997. GUYANA: Potaro River, 31 Dec 1948, Atkinson 85; Mt. Kainama, Potaro District, 1600 ft, 20 Oct 1959, Whitton 122; Upper Mazaruni River Basin, Merume Mountain, trail along SE ridge, 1140 m alt, 4-8 Jul 1960, Tillett et al 43995. SURINAM: Bakhuis mountains, from camp 3 SW to top, 840 m alt, 24 Feb 1965, Maas 3005. BRAZIL: Road to Cerro de la Neblina from Rio Tucano, 1250 m alt, 21 Apr 1964, Ewel 123, 138; Cerro de la Neblina, between Missao and Serra Pirapucu, catinga forest, 400-800 m alt, 13 Jan 1966, Silva & Brazdo 60813.
Distribution and Ecology - Distribution. VENEZUELA. Edo. Bolivar: Cerro Auyan-tepui, Alto Caroni, 1600 m alt, Jan 1949, Cardona 2635; Ptari-tepui, vicinity of Cave Rock, 1600-2000 m alt, 14-19 Aug 1970, Moore et al 9734; Chimanta Massif, Central Section, vicinity of Upper Falls of Rio Tirica above Summit Camp, 1940-1950 m alt, 7 Feb 1955, Steyermark & Wurdack 519, 553; Steyermark Falls on upper reaches of Rio Tirica, 1700-1750 m alt, 3 Jul 1953, Steyermark 76012; Apacara-tepui, between Camp 3 and main line of sandstone bluffs, 1210-1600 m alt, 17 Apr 1953, Steyermark 75086; Chimanta-tepui (Torono-tepuí), above valley of Rio Tirica, 1000-1700 m alt, 16 May 1953, Steyermark 75454; Gran Sabana, Ilu-tepui, Camp 1, 1000 m alt, 10 Mar 1952, Maguire 33334; Rio Aponguao drainage ca km 146 south of El Dorado, 1350-1400 m alt, 21 Dec 1970, Steyermark et al 104214; Cerro Venamo, southwest part near Rio Venamo, 950-1150 m alt, 29-30 Dec 1963, Steyermark et al 92404; Cerro Venamo, northwest slopes, 1100-1300 m alt, 14 Apr 1960, Steyermark & Nilsson 134; Cerro Venamo region, between campamento 125 and km 127, wooded ridge of Fila de La Danta, 1200 m alt, 15-17 Apr 1960, Steyermark & Nilsson 207; Meseta del Jaua, Cerro Jaua, southwest portion, along tributary of Rio Marajano, 1750-1800 m alt, 22-28 Feb 1974, Steyermark et al 109659; Cerro Guanacoco, northwest portion, 1450 m alt, 3 Mar 1974, Steyermark et al 109756; Salto Angel, 1200 m alt, 18 Aug 1968, Foldats 7211; slopes of Cerro Uei, 860-1050 m alt, 27-28 Dec 1970, Steyermark et al 104560; Rio Cuyuni drainage, at km 142-143 S of El Dorado, 1300-1380 m alt, 22-28 Dec 1970, Steyermark et al 104260; Sierra W of waterfall of Rio Chixanan, 80 km southeast of El Dorado, 700 m alt, 29 Aug 1961, Steyermark 89608; Sierra Pacaraima, headwaters of Rio Paragua, 1400 m alt, 4-5 Mar 1973, Steyermark 107232; Cerro Guaiquinima, Alto Rio Paragua, 300-500 m alt, Cardona 901, 1077; slopes of Cerro Opacara, 900 m alt, 26 Jun 1946, Cardona 1601. Territorio de Amazonas: Cerro Huachamacari, Rio Cunucunuma, base of East Ridge, 1500 m alt, 8 Dec 1950, Maguire et al 30044; Cerro Marahuaca, 1600 m alt, 10 May 1949, Maguire & Maguire 29174; Cerro de la Neblina, just S of Camp 3, 650-700 m alt, 23 Dec 1953, aguire et al. 36812; Cerro Yapacana, 825 m alt, 4 May 1970, Steyermark & Bunting 103086; Serrania Yutaja, Rio Manapiare, Northwest Ridge, 1400 m alt, 11 Feb 1953, Maguire & Maguire 35160; Cerro Yutaje, left fork of Caño Yutaje, 1250 m alt, 12 Feb 1953, Maguire & Maguire 35201; Serrania Paru, around Laguna Asisa, Cerro Asisa, 1310 m alt, 7 May 1973, Hoyos &. Morillo 54; region of Duida and Marahuaca, Campo Benitez, Apr-Jun 1950, Barnes 55572; Cerro Sipapo, Forest Phelps Camp to North Savanna, 17 Dec 1948, Maguire & Politi 27753; Cerro Duida, Culebra Creek Basin, 1400 m alt, 23 Apr 1949, Maguire & Maguire 29143; Cerro Duida, near escarpment at 1400 m alt, 23 Apr 1949, Maguire & Maguire 29145; Cerro Duida, ridge W of Caño Culebra, 1700 m alt, 22 Nov 1950, Maguire et al 29707; Cerro de la Neblina, Rio Yatua, trail between Camp 2 and 3, 500-700 m alt, 7 Nov 1957, Maguire et al 41997. GUYANA: Potaro River, 31 Dec 1948, Atkinson 85; Mt. Kainama, Potaro District, 1600 ft, 20 Oct 1959, Whitton 122; Upper Mazaruni River Basin, Merume Mountain, trail along SE ridge, 1140 m alt, 4-8 Jul 1960, Tillett et al 43995. SURINAM: Bakhuis mountains, from camp 3 SW to top, 840 m alt, 24 Feb 1965, Maas 3005. BRAZIL: Road to Cerro de la Neblina from Rio Tucano, 1250 m alt, 21 Apr 1964, Ewel 123, 138; Cerro de la Neblina, between Missao and Serra Pirapucu, catinga forest, 400-800 m alt, 13 Jan 1966, Silva & Brazdo 60813.
Discussion:
1. Cavendishia duidae A. C. Smith, Bull. Torrey Club 58: 443. 1931. Cavendishia gleasoniana A. C. Smith, Bull. Torrey Club 60: 117. 1933. Cavendishia phelpsiae Camp, Brillonia 7: 85. 1950. Cavendishia gleasoniana was distinguished by A. C. Smith from C. duidae on the basis of its proportionately broader, bullate leaves and shorter, fewer-flowered inflorescences. Cavendishia phelpsiae was distinguished by Camp from C. duidae on the basis of its smaller and differently shaped leaves, and its pubescent twigs. In the suite of material now available, these differences no longer can be applied to separate these taxa. All agree in having the inner surface of the corolla tube glabrous to sparsely pilose, although externally glabrous, the inner surface of the filaments pilose above the base and towards the middle, whereas the dorsal external face is glabrous. All have the filaments and anthers of shorter and longer stamens of similar lengths, including the lengths of the tubules and anther sacs. inflorescence, at first thought to distinguish C. duidae by its more elongate type with a greater number of flowers, cannot be rehed upon as a character, nor can the supposedly fewer-flowered, shorter inflorescence of C. gleasoniana be used as an infallible character. Many specimens combine the broad ovate leaf with bullate upper surface (supposedly characteristic of C. gleasoniana) with the more elongated type of inflorescence (supposedly found in C. duidae). Conversely, the more elongated, ovate-lanceolate or oblong-lanceolate leaf may be combined with the shorter, denser inflorescence. Likewise, the pubescence of the lower surface of the leaf blades varies in the specimens at hand from completely glabrous to hispid on the main nerves beneath, and from the pubescent lower surface described originally for C. gleasoniana to one with the lower surface glabrous. The stems, likewise vary from glabrous to strongly pilose-hispid. Often juvenile twigs are densely setose-pilose while the older branches on the same plant are glabrous. A similar situation was found in Central America by Luteyn (1976) in a group of three species (which are in turn closely related to C. duidae sl). There too, the characters of leaf size and shape, the degree of bullateness and pubescence, and twig pubescence were found to vary sporadically throughout the range of the species, and without any correlation with geography or morphology.
1. Cavendishia duidae A. C. Smith, Bull. Torrey Club 58: 443. 1931. Cavendishia gleasoniana A. C. Smith, Bull. Torrey Club 60: 117. 1933. Cavendishia phelpsiae Camp, Brillonia 7: 85. 1950. Cavendishia gleasoniana was distinguished by A. C. Smith from C. duidae on the basis of its proportionately broader, bullate leaves and shorter, fewer-flowered inflorescences. Cavendishia phelpsiae was distinguished by Camp from C. duidae on the basis of its smaller and differently shaped leaves, and its pubescent twigs. In the suite of material now available, these differences no longer can be applied to separate these taxa. All agree in having the inner surface of the corolla tube glabrous to sparsely pilose, although externally glabrous, the inner surface of the filaments pilose above the base and towards the middle, whereas the dorsal external face is glabrous. All have the filaments and anthers of shorter and longer stamens of similar lengths, including the lengths of the tubules and anther sacs. inflorescence, at first thought to distinguish C. duidae by its more elongate type with a greater number of flowers, cannot be rehed upon as a character, nor can the supposedly fewer-flowered, shorter inflorescence of C. gleasoniana be used as an infallible character. Many specimens combine the broad ovate leaf with bullate upper surface (supposedly characteristic of C. gleasoniana) with the more elongated type of inflorescence (supposedly found in C. duidae). Conversely, the more elongated, ovate-lanceolate or oblong-lanceolate leaf may be combined with the shorter, denser inflorescence. Likewise, the pubescence of the lower surface of the leaf blades varies in the specimens at hand from completely glabrous to hispid on the main nerves beneath, and from the pubescent lower surface described originally for C. gleasoniana to one with the lower surface glabrous. The stems, likewise vary from glabrous to strongly pilose-hispid. Often juvenile twigs are densely setose-pilose while the older branches on the same plant are glabrous. A similar situation was found in Central America by Luteyn (1976) in a group of three species (which are in turn closely related to C. duidae sl). There too, the characters of leaf size and shape, the degree of bullateness and pubescence, and twig pubescence were found to vary sporadically throughout the range of the species, and without any correlation with geography or morphology.
Distribution:
Venezuela South America| Guyana South America| Suriname South America|
Venezuela South America| Guyana South America| Suriname South America|