Monographs Details:
Authority:
Luteyn, James L. 1976. A revision of the Mexican Central American species of Cavendishia (Vacciniaceae). Mem. New York Bot. Gard. 28 (3): 1-138.
Luteyn, James L. 1976. A revision of the Mexican Central American species of Cavendishia (Vacciniaceae). Mem. New York Bot. Gard. 28 (3): 1-138.
Family:
Ericaceae
Ericaceae
Description:
Species Description - Erect, often arching, terrestrial shrub, 2.5-3 m tall; stem base about 2.5 cm in diameter. Mature branches terete, glabrous, about 1 cm in diameter; bark yellowish-brown; immature branches and twigs of new growth bluntly angled or ribbed, minutely striate, glabrous. Petioles terete, sHghtiy flattened adaxially, rugose (2.5-)6-8(-10) mm long and 2-2.5(-3.5) mm in diameter, glabrous or infrequently puberulent, golden-tan. Leaves elhptic, lance-elliptic to ovatelanceolate, 9-16 cm long and 3-6 cm broad, basally acute to narrowly obtuse, rarely rounded, apically acuminate to somewhat caudate-acimiinate, olive-green to grayish-green above and tannish-green beneath when dry, glabrous, but with scattered, elevated, reddish globular glands, 0.1-0.2 mm in diameter along upper leaf surface, these often caducous leaving a reddish or blackish papillate scar; 5(-7)-plinerved, innermost lateral nerves often arising 2-3 cm above base, midrib impressed above, raised and conspicuous beneath, lateral nerves impressed above becoming plane to slightiy elevated distally with outer lateral nerves often obscure beyond middle, raised and conspicuous beneath, veinlets raised and usuaUy conspicuous above, obscure to slightly raised beneath. Infiorescence viscid, 11-21(-26) flowered (often with lowest several nodes Sterile); rachis flattened, gradually tapering and often flexuous distally, conspicuously ribbed and minutely striate, glabrous or rarely puberulent, (3-)8-15 cm long and up to 5 mm in diameter at base, pink to dark-pink when fresh, provided along its length and especially at base with tiny, peltate, globular reddish glands. Floral bracts glabrous, oblong to oblanceolate, rarely obovate, basally narrowed to a slightly clasping base, apically rounded to narrowly obtuse, 3-5 cm long and 1-2 cm broad, pink, marginaUy with 7-23 dark red, globular, peltate or sessile glands about 0.7 mm in diameter. Pedicels subterete, swollen distally, striate, glabrous, (11 -) 15-19 mm long and 1-1.5 mm in diameter, pink, with few, scattered, globular, reddish glands. Bracteoles ovate to lanceolate, I.5-2(-4) mm long and 1-2 mm broad, pink, with few, scattered, dark red, globular glands both along margins and apically. Calyx glabrous, 4-6 mm long; tube cylindric or spreading, rugose to strongly ribbed, 1.5-2 mm long and 2.5 mm in diameter, nonapophysate, green, usually so densely covered by dark red, globular to disc-shaped, peltate glands as to obscure surface; limb cylindriccampanulate, smooth, (2.5-)3-3.5 mm long including lobes, pale-green to pinkish, with scattered, globular to disc-shaped, peltate glands; lobes triangular, obtuse, 1-1.5 mm long and 1.5 mm broad, erect after anthesis, marginally glandular callose-thickened with glands contiguous at lobe apices, pinkish-green with dark pink margins; sinus broadly rounded. Corolla constricted basally, narrowed to thorat, 2.5-3.5 cm long and 8-10 mm in diameter, glabrous or pilose (then glabrous at constricted base), white; lobes triangular, obtuse, 1-2 mm long. Stamens 27-30 mm long; filaments slightly coherent basally, strigose distally, alternately either 4-6 mm or 10-13.5 mm long, white; anthers including tubules alternately either 17-20.5 mm or 25-27 mm long, yellow; thecae 7.5-11 mm long. Style 25-31 mm long. Berry not seen. Phenology: Flowering January-May.
Species Description - Erect, often arching, terrestrial shrub, 2.5-3 m tall; stem base about 2.5 cm in diameter. Mature branches terete, glabrous, about 1 cm in diameter; bark yellowish-brown; immature branches and twigs of new growth bluntly angled or ribbed, minutely striate, glabrous. Petioles terete, sHghtiy flattened adaxially, rugose (2.5-)6-8(-10) mm long and 2-2.5(-3.5) mm in diameter, glabrous or infrequently puberulent, golden-tan. Leaves elhptic, lance-elliptic to ovatelanceolate, 9-16 cm long and 3-6 cm broad, basally acute to narrowly obtuse, rarely rounded, apically acuminate to somewhat caudate-acimiinate, olive-green to grayish-green above and tannish-green beneath when dry, glabrous, but with scattered, elevated, reddish globular glands, 0.1-0.2 mm in diameter along upper leaf surface, these often caducous leaving a reddish or blackish papillate scar; 5(-7)-plinerved, innermost lateral nerves often arising 2-3 cm above base, midrib impressed above, raised and conspicuous beneath, lateral nerves impressed above becoming plane to slightiy elevated distally with outer lateral nerves often obscure beyond middle, raised and conspicuous beneath, veinlets raised and usuaUy conspicuous above, obscure to slightly raised beneath. Infiorescence viscid, 11-21(-26) flowered (often with lowest several nodes Sterile); rachis flattened, gradually tapering and often flexuous distally, conspicuously ribbed and minutely striate, glabrous or rarely puberulent, (3-)8-15 cm long and up to 5 mm in diameter at base, pink to dark-pink when fresh, provided along its length and especially at base with tiny, peltate, globular reddish glands. Floral bracts glabrous, oblong to oblanceolate, rarely obovate, basally narrowed to a slightly clasping base, apically rounded to narrowly obtuse, 3-5 cm long and 1-2 cm broad, pink, marginaUy with 7-23 dark red, globular, peltate or sessile glands about 0.7 mm in diameter. Pedicels subterete, swollen distally, striate, glabrous, (11 -) 15-19 mm long and 1-1.5 mm in diameter, pink, with few, scattered, globular, reddish glands. Bracteoles ovate to lanceolate, I.5-2(-4) mm long and 1-2 mm broad, pink, with few, scattered, dark red, globular glands both along margins and apically. Calyx glabrous, 4-6 mm long; tube cylindric or spreading, rugose to strongly ribbed, 1.5-2 mm long and 2.5 mm in diameter, nonapophysate, green, usually so densely covered by dark red, globular to disc-shaped, peltate glands as to obscure surface; limb cylindriccampanulate, smooth, (2.5-)3-3.5 mm long including lobes, pale-green to pinkish, with scattered, globular to disc-shaped, peltate glands; lobes triangular, obtuse, 1-1.5 mm long and 1.5 mm broad, erect after anthesis, marginally glandular callose-thickened with glands contiguous at lobe apices, pinkish-green with dark pink margins; sinus broadly rounded. Corolla constricted basally, narrowed to thorat, 2.5-3.5 cm long and 8-10 mm in diameter, glabrous or pilose (then glabrous at constricted base), white; lobes triangular, obtuse, 1-2 mm long. Stamens 27-30 mm long; filaments slightly coherent basally, strigose distally, alternately either 4-6 mm or 10-13.5 mm long, white; anthers including tubules alternately either 17-20.5 mm or 25-27 mm long, yellow; thecae 7.5-11 mm long. Style 25-31 mm long. Berry not seen. Phenology: Flowering January-May.
Discussion:
Cavendishia panamensis is a distinctive species characterized at once by profusely glandular-margined floral bracts and bracteoles, and densely glandcovered calyx tubes (so densely so as to obscure the surface from view). In these respects it is reminiscent of C. chiriquiensis and C. pseudostenophylla. Some of the more salient differences between these three taxa have been compared in Table VI in the discussion of C. pseudostenophylla. Cavendishia panamensis also differs from C. chiriquiensis by a basic difference in calyx lobe glands. In C. panamensis the callose-thickened portion of the calyx lobe is strictiy marginal, whne in C. chiriquiensis the entire calyx lobe is callose-thickened and appears occasionally to break off.
Panama. Province of Chiriqui. Boquete District, Bajo Chorro Trail, 2160 m, Luteyn 3056 (Holotype: DUKE! isotypes: F! GH! LL! MICH! MO! NY! PMA! US!).
Distribution:
Panama Central America|
Panama Central America|