Monographs Details:
Authority:
Luteyn, James L. 1983. Ericaceae--part I. Cavendishia. Fl. Neotrop. Monogr. 35: 1-290. (Published by NYBG Press)
Luteyn, James L. 1983. Ericaceae--part I. Cavendishia. Fl. Neotrop. Monogr. 35: 1-290. (Published by NYBG Press)
Family:
Ericaceae
Ericaceae
Synonyms:
Thibaudia punctata Ruiz & Pav. ex J.St.-Hil., Thibaudia punctatifolia Ruiz & Pav. ex G.Don, Cavendishia punctatifolia (Ruiz & Pav. ex G.Don) Hoerold, Cavendishia kraenzliniana Hoerold
Thibaudia punctata Ruiz & Pav. ex J.St.-Hil., Thibaudia punctatifolia Ruiz & Pav. ex G.Don, Cavendishia punctatifolia (Ruiz & Pav. ex G.Don) Hoerold, Cavendishia kraenzliniana Hoerold
Description:
Species Description - Terrestrial shrub usually with one main branch arching to 4 m; mature branches subterete and often appearing hollow after drying, striate, glabrous, brownish; twigs pruinose. Leaves oblong or lance-elliptic, 12-20(-26) X 4-8.5(-11) cm, basally rounded or broadly obtuse, apically acute or short-acuminate, glabrous, conspicuously punctate on both surfaces after recessed glandular fimbriae break off; 3-5(-7)-plinerved with inner pair of lateral nerves arising 1.5-2.5 cm above base, midrib, lateral and reticulate veins impressed above and raised beneath; axillary bud scales rarely appearing pseudostipular; petioles subterete, adaxially flattened, rugose, glabrous, 5-14 mm long and 3-4 mm diam. Inflorescence racemose (rarely with one or two secondary branches near base), 30-40(-60)-flow-ered, encircled at base by numerous, subcoriaceous, ovate, apiculate, glandular-fimbriate-margined bracts to 15 mm long and broad, these often persistent through anthesis; rachis subterete, usually conspicuously striate or ridged, glabrous, 1030 cm long and 3-5 mm in diam., secondary branches 4-7(-11) cm long when present; floral bracts caducous, oblong, 8-15 X 5-7 mm, marginally glandular-fimbriate, green suffused with red when fresh; pedicels subterete, ridged and angled, glabrous, 10-12 mm long at anthesis but elongating to 36 mm in fruit; bracteoles subpersistent, broadly to narrowly ovate, acute to long-acuminate, 1-2.8 mm long, marginally glandular-fimbriate. Flowers: calyx glabrous, 5-6 mm long, green suffused with red; hypanthium globose to subcylindric, rugose, 2.2-2.5 mm long, basally slightly apophysate; limb spreading, rugose, 2.5-3.5 mm long; lobes oblong-ovate to hemispheric, apiculate, 1-2 X 2.6 mm, erect after anthesis, entire margin thin lacerate-glandular and reddish in color; sinus acute; corolla subcylindric to conical-cylindric tapering distally, glabrous, 10-15 mm long, pale yellowish-green when fresh, lobes triangular, acute, ca. 1 mm long; stamens 8.7-10.3 mm; filaments densely short-pilose ventrally, alternately 2.2-2.8 mm and 3-3.5 mm long; anthers alternately 7.2-8.6 mm and 7-8.3 mm long; thecae alternately 2.8-3 mm and 3-4 mm long; style equaling corolla. Young berry ca. 12 mm diam., dark wine-red when fresh.
Species Description - Terrestrial shrub usually with one main branch arching to 4 m; mature branches subterete and often appearing hollow after drying, striate, glabrous, brownish; twigs pruinose. Leaves oblong or lance-elliptic, 12-20(-26) X 4-8.5(-11) cm, basally rounded or broadly obtuse, apically acute or short-acuminate, glabrous, conspicuously punctate on both surfaces after recessed glandular fimbriae break off; 3-5(-7)-plinerved with inner pair of lateral nerves arising 1.5-2.5 cm above base, midrib, lateral and reticulate veins impressed above and raised beneath; axillary bud scales rarely appearing pseudostipular; petioles subterete, adaxially flattened, rugose, glabrous, 5-14 mm long and 3-4 mm diam. Inflorescence racemose (rarely with one or two secondary branches near base), 30-40(-60)-flow-ered, encircled at base by numerous, subcoriaceous, ovate, apiculate, glandular-fimbriate-margined bracts to 15 mm long and broad, these often persistent through anthesis; rachis subterete, usually conspicuously striate or ridged, glabrous, 1030 cm long and 3-5 mm in diam., secondary branches 4-7(-11) cm long when present; floral bracts caducous, oblong, 8-15 X 5-7 mm, marginally glandular-fimbriate, green suffused with red when fresh; pedicels subterete, ridged and angled, glabrous, 10-12 mm long at anthesis but elongating to 36 mm in fruit; bracteoles subpersistent, broadly to narrowly ovate, acute to long-acuminate, 1-2.8 mm long, marginally glandular-fimbriate. Flowers: calyx glabrous, 5-6 mm long, green suffused with red; hypanthium globose to subcylindric, rugose, 2.2-2.5 mm long, basally slightly apophysate; limb spreading, rugose, 2.5-3.5 mm long; lobes oblong-ovate to hemispheric, apiculate, 1-2 X 2.6 mm, erect after anthesis, entire margin thin lacerate-glandular and reddish in color; sinus acute; corolla subcylindric to conical-cylindric tapering distally, glabrous, 10-15 mm long, pale yellowish-green when fresh, lobes triangular, acute, ca. 1 mm long; stamens 8.7-10.3 mm; filaments densely short-pilose ventrally, alternately 2.2-2.8 mm and 3-3.5 mm long; anthers alternately 7.2-8.6 mm and 7-8.3 mm long; thecae alternately 2.8-3 mm and 3-4 mm long; style equaling corolla. Young berry ca. 12 mm diam., dark wine-red when fresh.
Discussion:
In the Herbarium Pavón (MA) the labels of the Ruiz & Pav6n collections bear the epithets “Thibaudia punctata” and “Thibaudia punctatifolia” In the manu script for vol. 4 of the Flora Peruviana et Chilensis (published in 1955), Ruiz and Pav6n described and illustrated Thibaudia punctatifolia as a new species “in Peruviae Andium nemoribus ad montium fauces in Huassahuassi, Lanco et Chu-rupallana tractibus et copiose in Panatahuarum Provincia ad Acomayo, Pillao et Mufia.” This manuscript, however, was not published in 1802 as anticipated although the plates were distributed (about this time, but exact date unknown) to several botanists by O. Rich thus providing effective publication (see Stafleu, 1967).The first Ruiz and Pavón Ericaceae to be validly published were Thibaudia mellifera (R. & P.), T. punctata (R. & P.), and T. bracteata (R. & P.), described by Jaume St.-Hilaire in 1805. Later, D. Don (1834) gave short diagnoses for some of Ruiz’s plants he found in Lambert’s herbarium. Don adapted some of the manuscript names of Ruiz but did not consider the names of St.-Hilaire. Thus Don validated T. punctatifolia (R. & P.) and was later followed by Hoerold (1909) who made the combination Cavendishia punctatifolia. In the same work Hoerold published C. kraenzliniana without comment as to its relationships.Smith (1932) followed Hoerold in the usage of C. punctatifolia stating that he [Smith] felt the names on the plates (of vol. 4, Fl. peruv. chilen.) were authentic. Furthermore, Smith (p. 481) listed the type as “collected by Ruiz and Pavón (or by Dombey?),” and cited the lectotype as “Dombev 431 (B, type collection?).” Smith had not seen material at MA, and in view of his stated doubts about the type and collectors, I feel justified in lectotypifying this species on the basis of the Ruiz and Pavón collection deposited at MA.Cavendishia punctata is more closely related to martii than the key may indicate. It differs from C. martii only by its usually racemose (not paniculate) inflorescence, oblong and conspicuously punctate (not ovate and less conspicuously punctate) leaves, and its distribution. Three collections of punctata, however, (Boeke 1156, Killip & Smith 24473, and Luteyn & Lebrdn-Luteyn 5427) have racemes with 1-4 short, weakly developed branches and in this respect approaching paniculate C. martii in this character. Additional collections from southern San Martin-northern Huanuco and from Huancavelica-southem Junin Depts. are sorely needed to better establish the relationships between these two species.
In the Herbarium Pavón (MA) the labels of the Ruiz & Pav6n collections bear the epithets “Thibaudia punctata” and “Thibaudia punctatifolia” In the manu script for vol. 4 of the Flora Peruviana et Chilensis (published in 1955), Ruiz and Pav6n described and illustrated Thibaudia punctatifolia as a new species “in Peruviae Andium nemoribus ad montium fauces in Huassahuassi, Lanco et Chu-rupallana tractibus et copiose in Panatahuarum Provincia ad Acomayo, Pillao et Mufia.” This manuscript, however, was not published in 1802 as anticipated although the plates were distributed (about this time, but exact date unknown) to several botanists by O. Rich thus providing effective publication (see Stafleu, 1967).The first Ruiz and Pavón Ericaceae to be validly published were Thibaudia mellifera (R. & P.), T. punctata (R. & P.), and T. bracteata (R. & P.), described by Jaume St.-Hilaire in 1805. Later, D. Don (1834) gave short diagnoses for some of Ruiz’s plants he found in Lambert’s herbarium. Don adapted some of the manuscript names of Ruiz but did not consider the names of St.-Hilaire. Thus Don validated T. punctatifolia (R. & P.) and was later followed by Hoerold (1909) who made the combination Cavendishia punctatifolia. In the same work Hoerold published C. kraenzliniana without comment as to its relationships.Smith (1932) followed Hoerold in the usage of C. punctatifolia stating that he [Smith] felt the names on the plates (of vol. 4, Fl. peruv. chilen.) were authentic. Furthermore, Smith (p. 481) listed the type as “collected by Ruiz and Pavón (or by Dombey?),” and cited the lectotype as “Dombev 431 (B, type collection?).” Smith had not seen material at MA, and in view of his stated doubts about the type and collectors, I feel justified in lectotypifying this species on the basis of the Ruiz and Pavón collection deposited at MA.Cavendishia punctata is more closely related to martii than the key may indicate. It differs from C. martii only by its usually racemose (not paniculate) inflorescence, oblong and conspicuously punctate (not ovate and less conspicuously punctate) leaves, and its distribution. Three collections of punctata, however, (Boeke 1156, Killip & Smith 24473, and Luteyn & Lebrdn-Luteyn 5427) have racemes with 1-4 short, weakly developed branches and in this respect approaching paniculate C. martii in this character. Additional collections from southern San Martin-northern Huanuco and from Huancavelica-southem Junin Depts. are sorely needed to better establish the relationships between these two species.
Distribution:
Peru South America| Huánuco Peru South America| Junín Peru South America|
Peru South America| Huánuco Peru South America| Junín Peru South America|
Common Names:
huehli del monte, huegli
huehli del monte, huegli