Monographs Details:
Authority:
Luteyn, James L. 1983. Ericaceae--part I. Cavendishia. Fl. Neotrop. Monogr. 35: 1-290. (Published by NYBG Press)
Luteyn, James L. 1983. Ericaceae--part I. Cavendishia. Fl. Neotrop. Monogr. 35: 1-290. (Published by NYBG Press)
Family:
Ericaceae
Ericaceae
Description:
Species Description - Epiphytic shrub, mature branches terete, striate, glabrate; twigs hispid with brownish hairs to 2.2 mm long. Leaves elliptic to ovate-elliptic, (5-) - 1-17 X (1.5-) 3.5-7 cm, basally rounded, apically abruptly short-caudate, sparsely hispid and glandular-fimbriate on both surfaces; 5(-7)-plinerved from base, midrib impressed above and raised beneath, inner pair of secondary nerves plane or slightly compressed above and raised beneath, remaining secondary nerves and reticulate veinlets slightly raised but inconspicuous on both surfaces; petioles terete, rugose, hispid, 4-8 mm long and 1-2 mm in diam. Inflorescence ca. 20-30-flowered, encircled at base by numerous, subcoriaceous, striate, oblong to ovate, emarginate bracts to 15 mm long; rachis subterete, sparsely hispid, ca. 8-10 cm long and ca. 3 mm in basal diam.; floral bracts membranaceous, venation reticulate and conspicuous, obovate, glandular-fimbriate on both surfaces, glabrous or ciliate, those bracts of the lower half of inflorescence subtending sterile nodes 30-35 X 25 mm, apically emarginate, those of the upper half similar but subtending fertile nodes and 15 mm broad, pale greenish-white; pedicels terete, glabrous, 5-5.5 mm long and ca. 1 mm in diam.; bracteoles membranaceous, oblanceolate, apically emarginate, ca. 8 mm long and 3 mm broad at distal tip, ciliate and also with scattered glandular fimbriae dorsally. Flowers: calyx glabrous (except the ciliate lobes), 30-33 mm long; hypanthium cylindric, 3-4 mm long; limb membranaceous, infundibuliform, 27-30 mm long, densely glandular-fimbriate, conspicuously veined, the venation free-ending; lobes narrowly imbricate, oblong-obovate, 13-14 mm long and 8-9 mm broad across the top, apically rounded, emarginate and obscurely callose-apiculate, ciliate or not; sinus acute; corolla cylindric, glabrous, 36-37 mm long and 7-8 mm in diam., lobes triangular, obtuse, ca. 1.5 mm long; stamens alternately 15-15.5 mm and 14-14.5 mm long; filaments glabrous, alternately 2 mm and 4.2-4.7 mm long; anthers alternately 13-13.3 mm and 11.5 mm long; thecae slightly granular, 4.6 mm long; tubules dehiscing by short, oval, oblique pores, 0.7-1.4 mm long; style ca. 36 mm long. Berry not seen.
Species Description - Epiphytic shrub, mature branches terete, striate, glabrate; twigs hispid with brownish hairs to 2.2 mm long. Leaves elliptic to ovate-elliptic, (5-) - 1-17 X (1.5-) 3.5-7 cm, basally rounded, apically abruptly short-caudate, sparsely hispid and glandular-fimbriate on both surfaces; 5(-7)-plinerved from base, midrib impressed above and raised beneath, inner pair of secondary nerves plane or slightly compressed above and raised beneath, remaining secondary nerves and reticulate veinlets slightly raised but inconspicuous on both surfaces; petioles terete, rugose, hispid, 4-8 mm long and 1-2 mm in diam. Inflorescence ca. 20-30-flowered, encircled at base by numerous, subcoriaceous, striate, oblong to ovate, emarginate bracts to 15 mm long; rachis subterete, sparsely hispid, ca. 8-10 cm long and ca. 3 mm in basal diam.; floral bracts membranaceous, venation reticulate and conspicuous, obovate, glandular-fimbriate on both surfaces, glabrous or ciliate, those bracts of the lower half of inflorescence subtending sterile nodes 30-35 X 25 mm, apically emarginate, those of the upper half similar but subtending fertile nodes and 15 mm broad, pale greenish-white; pedicels terete, glabrous, 5-5.5 mm long and ca. 1 mm in diam.; bracteoles membranaceous, oblanceolate, apically emarginate, ca. 8 mm long and 3 mm broad at distal tip, ciliate and also with scattered glandular fimbriae dorsally. Flowers: calyx glabrous (except the ciliate lobes), 30-33 mm long; hypanthium cylindric, 3-4 mm long; limb membranaceous, infundibuliform, 27-30 mm long, densely glandular-fimbriate, conspicuously veined, the venation free-ending; lobes narrowly imbricate, oblong-obovate, 13-14 mm long and 8-9 mm broad across the top, apically rounded, emarginate and obscurely callose-apiculate, ciliate or not; sinus acute; corolla cylindric, glabrous, 36-37 mm long and 7-8 mm in diam., lobes triangular, obtuse, ca. 1.5 mm long; stamens alternately 15-15.5 mm and 14-14.5 mm long; filaments glabrous, alternately 2 mm and 4.2-4.7 mm long; anthers alternately 13-13.3 mm and 11.5 mm long; thecae slightly granular, 4.6 mm long; tubules dehiscing by short, oval, oblique pores, 0.7-1.4 mm long; style ca. 36 mm long. Berry not seen.
Discussion:
Cavendishia chlamydantha is characterized by its large, membranaceous, infundibuliform calyx limb with narrowly imbricate lobes, and by the short, oval, oblique pores of its anthers. The margins of the floral bracts and calyx lobes are ciliate in the holotype (GH), while they are glabrous in the F isotype.In 1946, Smith noted that two other species named by him, C. micayensis and C. venosa, provided transitional features between C. chlamydantha and the more typical cavendishias. Although C. micayensis had short dehiscence pores like C. chlamydantha, its calyx limb was only somewhat infundibuliform and shorter (10-15 mm not 27-30 mm long) and its calyx lobes were of typical triangular shape. Cavendishia venosa, in contrast, had elongate dehiscence clefts typical for most species of Cavendishia, and its calyx was “normal” for the genus. I agree that C. micayensis is more closely related to C. chlamydantha and that C. venosa is further removed towards the more typical cavendishias; however, the available specimens of all members of subgen. Chlamydantha are so few that any statements concerning close relationships between species are speculative.
Cavendishia chlamydantha is characterized by its large, membranaceous, infundibuliform calyx limb with narrowly imbricate lobes, and by the short, oval, oblique pores of its anthers. The margins of the floral bracts and calyx lobes are ciliate in the holotype (GH), while they are glabrous in the F isotype.In 1946, Smith noted that two other species named by him, C. micayensis and C. venosa, provided transitional features between C. chlamydantha and the more typical cavendishias. Although C. micayensis had short dehiscence pores like C. chlamydantha, its calyx limb was only somewhat infundibuliform and shorter (10-15 mm not 27-30 mm long) and its calyx lobes were of typical triangular shape. Cavendishia venosa, in contrast, had elongate dehiscence clefts typical for most species of Cavendishia, and its calyx was “normal” for the genus. I agree that C. micayensis is more closely related to C. chlamydantha and that C. venosa is further removed towards the more typical cavendishias; however, the available specimens of all members of subgen. Chlamydantha are so few that any statements concerning close relationships between species are speculative.
Distribution:
Valle Colombia South America| Colombia South America|
Valle Colombia South America| Colombia South America|
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