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Family:

Ericaceae (Magnoliophyta)

Primary Citation:

Repert. Spec. Nov. Regni Veg. 22: 46. 1925

Common Names:

clavellina

Description:

Description: Evergreen shrub to ca. 3.5 m tall, with longitudinally furrowed bark; twigs slightly to moderately angled, slender, sparsely to densely lepidote, otherwise glabrous to densely pubescent; buds ovoid, 0.6-1 x 0.5-0.8 mm. Leaf blades widely obovate or orbicular, obovate to elliptic or narrowly elliptic, 1-5(-5.5) x 0.3-4.8 cm, ± flat to slightly or strongly recurved, coriaceous, ca. 0.3-0.48 mm thick; base narrowly to broadly cuneate to rounded; apex acute to rounded or truncate; margin plane to strongly revolute, entire to sinuous; venation brochidodromous, 3o veins ± reticulate; adaxial surface roughened by raised-reticulate pattern of 3o veins and/or thickened persistent scale bases (sometimes only clearly developed near margins or apex of leaf), to smooth (in var. toaensis), lepidote but scales quickly deciduous, sparsely to densely pubescent on basal portion of midvein, sometimes extending to some 2o and 3o veins, the 3o and higher order veins forming ± raised reticulum to nearly obscure, the midvein depressed or not, the 2o veins visible to obscure, not depressed or with few per leaf slighlty to strongly depressed; abaxial surface moderately lepidote, otherwise densely pubescent, the 3o and higher order veins ± flat, obscure to less commonly very slightly and laxly reticulate, the 2o veins slighlty raised and visible or obscure; scales rust colored, often becoming whitish with age, persistent, ca. 0.1-0.22 mm in diameter, ± irregular; petiole 1.5-6 mm long, lepidote, otherwise pubescent adaxially or all around; flower buds ± intermixed with vegetative buds. Inflorescences fasciculate, 1- to 7-flowered; pedicels not articulated with calyx and appearing swollen just below capsule, slender, 1.5-7 mm long, lepidote, otherwise glabrous to moderately pubescent; bracteoles subopposite, nearly basal, narrowly triangular, 0.5-0.8 mm long; floral bracts to ca. 0.8 mm long. Flowers 4-(or -5)-merous; calyx lobes triangular, with acuminate apices, 0.5-1 x 0.4-0.6 mm, adaxial side glabrous to very sparsely pubescent, especially toward margins or apex, abaxial side moderately to densely lepidote; corolla urceolate to cylindrical, white, 2-3 x 1-2.2 mm, abaxially sparsely lepidote; filaments roughened, 1-1.5 mm long, unappendaged or with very minute spurs near anther-filament junction; anthers 0.7-0.9 mm long; ovary lepidote, otherwise pubescent, placentae ± subapical. Capsules narrowly ovoid, 2.5-4.5 x 1-2 mm, lepidote, otherwise very sparsely pubescent (only a few hairs near base), the pale very thick sutures separating as unit from adjacent valves; seeds 1.5-2.4 mm long.

Distribution: Endemic to Cuba (Prov. Oriente, Sierra de Nipe, Sierra de Cristal, Sierra de Micara, Sierra del Frijol (Pico Galano), Sierra de Moa, and limonite plateau near Moa), in dense thickets, or moist montane forests, stream sides, grassy areas, Pinus cubensis savannas, at ca. 300-1200 m alt.

Key to the Varieties of Lyonia glandulosa: 1. Leaves flat, the veins not depressed, the lamina orbicular to broadly obovate, length/width quotient 0.9-1.8, apex always rounded ............. var. toaensis 1. Leaves slightly to strongly recurved, at least the midvein depressed, the lamina obovate to elliptic or narrowly elliptic, length/width quotient 1.5-7, apex acute to rounded. 2. Leaves obovate to less commonly elliptic, 0.5-2 cm wide, from 1.5-3(-3.5) times as long as wide, apical margin usually plane or only slightly revolute; Sierra de Nipe ............ var. glandulosa 2. Leaves elliptic to narrowly so, usually not over ca. 1 cm wide, from 2.5-7(-10) times as long as wide, apical margin slightly to strongly revolute; chiefly in Sierra de Micara, Sierra de Cristal, Sierra de Moa, and vicinity of Moa ............. var. revolutifolia