Taxon Details: Gaultheria lanigera Hook.f.
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Family:
Ericaceae (Magnoliophyta)
Ericaceae (Magnoliophyta)
Scientific Name:
Gaultheria lanigera Hook.f.
Gaultheria lanigera Hook.f.
Primary Citation:
Hooker's Icon. Pl. t. 66. 1837
Hooker's Icon. Pl. t. 66. 1837
Accepted Name:
This name is currently accepted.
This name is currently accepted.
Description:
Description: Erect, rhizomatous, shrub, 0.1-0.5(-1) m tall, occurring in large, straggling colonies; mature stems subterete, glabrate; bark grayish, cracking irregularly into thin strips; twigs subterete, complanate, densely lanate-villous with canescent to ferruginous, eglandular hairs to 6 mm long; buds ovate, complanate, scales reddish-brown, striate, glabrate. Leaves rigidly coriaceous, sparsely to densely scabrous (pustular) above due to scattered minute papillae, ovate, elliptic to subrotund, conspicuously concave or strongly revolute lengthwise, 1-2.5(-3.5) x 0.4-2(-2.8) cm, base rounded, subcordate to cordate, apex acute to rounded (bluntly mucronate), entire but margin strongly revolute, glabrous to weakly short-pilose above (lanate but becoming glabrous in bud), densely tomentose-lanate beneath (obscuring surface) with ferruginous to chocolate-brown, eglandular hairs; midrib and lateral nerves (ca. 3 per side) sometimes impressed above causing leaf surface to be bullate, raised but obscured by hairs beneath, reticulate veinlets plane to slightly raised or impressed above and obscure beneath; petiole subterete, flattened above, 2-9 mm long, densely villous-lanate, often glabrate. Inflorescence racemose, 2-10(-18)-flowered, bracteate basally; rachis subterete, bluntly angled, 1-8 cm long, sparsely white puberulent and also densely tomentose-villous-lanate with reddish to ferruginous hairs; pedicels subterete, 5-12 mm long, pubescent as rachis; bracteoles basal or located along basal 1/3 of pedicel, subcoriaceous to membranous, somewhat cochleariform, ovate to linear-ovate, 3-6 mm long, acute to acuminate, dorsally strigose-lanate, glabrate, ventrally glabrous to pilose; floral bracts subcoriaceous, cochleariform, striate, ovate-oblong, 4-8(-10) x 4-6 mm, acute to acuminate, pubescent as bracteoles. Flowers with calyx 6-8 mm long, lobes ovate-triangular, 5-6 x 2.3-3 mm, acuminate, ciliolate, dorsally tomentose-villous-lanate, ventrally puberulent to glabrous; corolla urceolate to cylindric-urceolate, 6-7.5(-9) x 4-6(-7) mm, tomentose-villous-lanate without, glabrous within, cream-colored to pale greenish-yellow or dark red, lobes ovate, obtuse or acute; stamens 5-5.5 mm long; filaments 3.2-4 mm long, densely pilose and also distally with a few rigid (swollen-headed) hairs along margins; anthers 1.7-2.5 mm long, awns conspicuous; ovary short-pilose or glabrous along sides and deciduously tomentose-lanate at apex; style 4-5 mm long, glabrous to short-pilose basally. Fruiting calyx 5-6 mm diam., blue-black.
Distribution: Rare in the Cordillera Oriental of north-central Colombia and then again in south-central Ecuador at elevations of (2800-)3000-3700 m. Flowering and fruiting throughout the season.
Type: Ecuador. Loja: Páramo between Oña and Saraguro, 3048 m, 1836 (fl), Jameson s.n. (holotype, K, photo NY s.n.).
Key to Varieties of Gaultheria lanigera:1. Lamina concave, not revolute lengthwise, (1-)1.5-2.5(-3.5) x (0.8-)1.2-2(-2.8) cm; petiole 3-9 mm long; inflorescence long-racemose, rachis 3-8 cm long, with 8-10(-18) flowers; floral bracts 6-8(-10) mm long; bracteoles located in basal 1/3 of pedicel, linear-ovate, 5-6 mm long; corolla cream-colored to pale greenish-yellow; Ecuador ......... var. lanigera
1. Lamina strongly revolute lengthwise, 1-2 x 0.4-0.6(-1.2) cm; petiole 2-3 mm long; inflorescence short-racemose, rachis about 1 cm long, with 2-4 flowers; floral bracts 4-5 mm long; bracteoles basal, ovate, ca. 3 mm long; corolla dark red; Colombia ..................... var. rufolanata
Description: Erect, rhizomatous, shrub, 0.1-0.5(-1) m tall, occurring in large, straggling colonies; mature stems subterete, glabrate; bark grayish, cracking irregularly into thin strips; twigs subterete, complanate, densely lanate-villous with canescent to ferruginous, eglandular hairs to 6 mm long; buds ovate, complanate, scales reddish-brown, striate, glabrate. Leaves rigidly coriaceous, sparsely to densely scabrous (pustular) above due to scattered minute papillae, ovate, elliptic to subrotund, conspicuously concave or strongly revolute lengthwise, 1-2.5(-3.5) x 0.4-2(-2.8) cm, base rounded, subcordate to cordate, apex acute to rounded (bluntly mucronate), entire but margin strongly revolute, glabrous to weakly short-pilose above (lanate but becoming glabrous in bud), densely tomentose-lanate beneath (obscuring surface) with ferruginous to chocolate-brown, eglandular hairs; midrib and lateral nerves (ca. 3 per side) sometimes impressed above causing leaf surface to be bullate, raised but obscured by hairs beneath, reticulate veinlets plane to slightly raised or impressed above and obscure beneath; petiole subterete, flattened above, 2-9 mm long, densely villous-lanate, often glabrate. Inflorescence racemose, 2-10(-18)-flowered, bracteate basally; rachis subterete, bluntly angled, 1-8 cm long, sparsely white puberulent and also densely tomentose-villous-lanate with reddish to ferruginous hairs; pedicels subterete, 5-12 mm long, pubescent as rachis; bracteoles basal or located along basal 1/3 of pedicel, subcoriaceous to membranous, somewhat cochleariform, ovate to linear-ovate, 3-6 mm long, acute to acuminate, dorsally strigose-lanate, glabrate, ventrally glabrous to pilose; floral bracts subcoriaceous, cochleariform, striate, ovate-oblong, 4-8(-10) x 4-6 mm, acute to acuminate, pubescent as bracteoles. Flowers with calyx 6-8 mm long, lobes ovate-triangular, 5-6 x 2.3-3 mm, acuminate, ciliolate, dorsally tomentose-villous-lanate, ventrally puberulent to glabrous; corolla urceolate to cylindric-urceolate, 6-7.5(-9) x 4-6(-7) mm, tomentose-villous-lanate without, glabrous within, cream-colored to pale greenish-yellow or dark red, lobes ovate, obtuse or acute; stamens 5-5.5 mm long; filaments 3.2-4 mm long, densely pilose and also distally with a few rigid (swollen-headed) hairs along margins; anthers 1.7-2.5 mm long, awns conspicuous; ovary short-pilose or glabrous along sides and deciduously tomentose-lanate at apex; style 4-5 mm long, glabrous to short-pilose basally. Fruiting calyx 5-6 mm diam., blue-black.
Distribution: Rare in the Cordillera Oriental of north-central Colombia and then again in south-central Ecuador at elevations of (2800-)3000-3700 m. Flowering and fruiting throughout the season.
Type: Ecuador. Loja: Páramo between Oña and Saraguro, 3048 m, 1836 (fl), Jameson s.n. (holotype, K, photo NY s.n.).
Key to Varieties of Gaultheria lanigera:
Flora and Monograph Treatment(s):
Gaultheria lanigera Hook.f.: [Article] Luteyn, James L., et al. 1995. Ericaceae, Part II. The Superior-Ovaried Genera (Monotropoideae, Pyroloideae, Rhododendroideae, and Vaccinioideae P.P.). Fl. Neotrop. Monogr. 66: 560.
Gaultheria lanigera Hook.f.: [Article] Luteyn, James L., et al. 1995. Ericaceae, Part II. The Superior-Ovaried Genera (Monotropoideae, Pyroloideae, Rhododendroideae, and Vaccinioideae P.P.). Fl. Neotrop. Monogr. 66: 560.