Taylor, W. Carl. 1976. Vascular flora of Jonca Creek, Ste Genevieve County, Missouri. Castanea. 41: 93-118.
Dryopteridaceae
Latin Diagnosis - Ab E. orbignyano (Fee) T. Moore atque E. plumoso (Fée) T. Moore squamis rhizomaticis pallide brunneis lanceolatis integris vel sparsim dentatis (non squamis rhizomaticis ciliatis) et in superficie laminari inferna punctis (non squamis) resinosis differt.
Species Description - Plants epiphytic and terrestrial; rhizomes compact, erect to very short-creeping, 4 mm in diameter, densely scaly, the scales lanceolate, 1.5-3.5 x 0.5-0.8 mm, pale reddish brown, entire to sparsely toothed; phyllopodia ca. 5 mm long; leaves approximate, 25- 45 x 1.7-3 cm; petioles 1/3-1/2 the sterile leaf length, stramineous to reddish brown, densely scaly, the scales ovate-lanceolate to lanceolate, to 4 x 1 mm, orange, patent, the margins ciliate, the cilia long, lax, pale, only occasionaly indurated cilia,; blades oblanceolate, chartaceous, apically acuminate, basally attenuate; veins evident, free, ca. I mm apart, set at ca. 700 to costae, hydathodes lacking but vein endings slightly thickened; blades adaxially moderately scaly, the scales ovate to lanceolate, long-ciliate, 0.5-1 x 0.3-0.7 mm, whitish, appressed, marginally with similar, but denser and slightly larger, spreading scales, abaxially with abundant yellowish to reddish, stalked resinour glands; fertile leaves about 1.5 times as long as the sterile ones, petioles ca. 2/3 the fertile leaf length, blades linear-oblanceolate, apically acuminate, basally long-decurrent, 15 × 0.5 cm; intersporangial scales lacking.
(Fig. 1 G-L)
This species is related to E. orbignyanum (F6e) T. Moore and E. plumosum (F6e) T. Moore, but differs from both in having pale brown, lanceolate, entire to sparsely toothed rhizome scales (vs. ciliate rhizome scales), and in bearing resinous dots (vs. scales) on the abaxial blade surface. It is further distinct from E. orbignyanum in having patent petiole scales with long, lax cilia (vs. appressed to spreading scales with short, stiff cilia) and from E. plumosum in the much longer petioles. The resinous dots on the abaxial blade would appear to indicate a relationship with E. mathewsii (Fée) T. Moore, but that species is very different in the overall scale pattern and probably not very closely related. In Elaphoglossum, resinous dots have apparently repeatedly evolved through the reduction of scales. Scale-dot intermediates can be observed in numerous species, such as in E. tenuiculum (Fée) Baker. We name this species for our friend and colleague, the Bolivian pteridologist Jasivia Gonzales.