Monographs Details:
Authority:
Mickel, John T. & Beitel, Joseph M. 1988. Pteridophyte Flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1-580.
Mickel, John T. & Beitel, Joseph M. 1988. Pteridophyte Flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1-580.
Family:
Dryopteridaceae
Dryopteridaceae
Description:
Species Description - Rhizome short- to moderately creeping, to 4 mm diam.; rhizome scales linear-lanceolate, dark brown, lustrous, subentire to only faintly toothed, ca. 5 mm long; phyllopodia distinct to indistinct; fronds clumped, 8-20 cm long, 0.8-1.2 cm wide; stipe 1/3-2/3 the frond length, with scales linear-lanceolate and dark-toothed or dark-tipped, spreading, 3 mm long, some more dissected like stellate hairs and appressed; blade narrowly elliptic, chartaceous, apex acute to acuminate, base narrowly cuneate; veins obscure, free, simple or once-forked, 0.6-1.0 mm apart, at 55-65° angle; hydathodes lacking; blade scales evenly and liberally distributed adaxially but not overlapping, linear-lanceolate, 1-1.5 mm long, with hair teeth, on lower surface often reduced to resinous dots; fertile fronds generally longer than the sterile and very long-stiped (ca. 5/6 the length), apex more acute or obtuse, base more broadly cuneate, scales on abaxial midvein but not among the sporangia; spores with slender, smooth crests with sparse spicules.
Species Description - Rhizome short- to moderately creeping, to 4 mm diam.; rhizome scales linear-lanceolate, dark brown, lustrous, subentire to only faintly toothed, ca. 5 mm long; phyllopodia distinct to indistinct; fronds clumped, 8-20 cm long, 0.8-1.2 cm wide; stipe 1/3-2/3 the frond length, with scales linear-lanceolate and dark-toothed or dark-tipped, spreading, 3 mm long, some more dissected like stellate hairs and appressed; blade narrowly elliptic, chartaceous, apex acute to acuminate, base narrowly cuneate; veins obscure, free, simple or once-forked, 0.6-1.0 mm apart, at 55-65° angle; hydathodes lacking; blade scales evenly and liberally distributed adaxially but not overlapping, linear-lanceolate, 1-1.5 mm long, with hair teeth, on lower surface often reduced to resinous dots; fertile fronds generally longer than the sterile and very long-stiped (ca. 5/6 the length), apex more acute or obtuse, base more broadly cuneate, scales on abaxial midvein but not among the sporangia; spores with slender, smooth crests with sparse spicules.
Discussion:
Acrostichum gratum Fée, Mém. foug. 8: 69. 1857. Syntypes. Mexico. México: Popocatépetl, Schajfner 279, 322b (P?). Acrostichum callolepis Fée, Mém. foug. 8: 69. 1857. Elaphoglossum callolepis (Fée) Moore, Index fil. 7. 1857. Type. Mexico. Galeotti s.n. (P?). Acrostichum roezlii Schaffner in Fée, Mém. foug. 8: 69. 1857. Elaphoglossum roezlii (Schaffner in Fée) Moore, Index fil. 14. 1857. Type. Mexico. México: Popocatépetl, Schaffner 280 (P?). Elaphoglossum gratum is closely allied to E. mathewsii and E. pringlei. It is somewhat like E. muelleri, which is also scaly, terrestrial and of high elevation, but can be distinguished by its linear-lanceolate, coarsely toothed blade scales, and E. muelleri has blade scales with little body to them and the rhizome scales are tan to orange. It also resembles E. petiolatum, which has entire rhizome scales, but E. gratum usually lacks the laminar resin dots. It is close to E. mathewsii in dark entire rhizome scales and heavier scaliness on top, and tendency for rhizome to creep, but E. mathewsii has a longer creeping rhizome and blade scales that are more deltoid-lanceolate and subentire, broader and with darker middle and pale margin rather than the usual white. Both may have the resin dots on the lower surface. The abaxial blade scales of E. gratum are generally somewhat folded and curved but not in ours, which are essentially glabrous beneath and may be distinct.
Acrostichum gratum Fée, Mém. foug. 8: 69. 1857. Syntypes. Mexico. México: Popocatépetl, Schajfner 279, 322b (P?). Acrostichum callolepis Fée, Mém. foug. 8: 69. 1857. Elaphoglossum callolepis (Fée) Moore, Index fil. 7. 1857. Type. Mexico. Galeotti s.n. (P?). Acrostichum roezlii Schaffner in Fée, Mém. foug. 8: 69. 1857. Elaphoglossum roezlii (Schaffner in Fée) Moore, Index fil. 14. 1857. Type. Mexico. México: Popocatépetl, Schaffner 280 (P?). Elaphoglossum gratum is closely allied to E. mathewsii and E. pringlei. It is somewhat like E. muelleri, which is also scaly, terrestrial and of high elevation, but can be distinguished by its linear-lanceolate, coarsely toothed blade scales, and E. muelleri has blade scales with little body to them and the rhizome scales are tan to orange. It also resembles E. petiolatum, which has entire rhizome scales, but E. gratum usually lacks the laminar resin dots. It is close to E. mathewsii in dark entire rhizome scales and heavier scaliness on top, and tendency for rhizome to creep, but E. mathewsii has a longer creeping rhizome and blade scales that are more deltoid-lanceolate and subentire, broader and with darker middle and pale margin rather than the usual white. Both may have the resin dots on the lower surface. The abaxial blade scales of E. gratum are generally somewhat folded and curved but not in ours, which are essentially glabrous beneath and may be distinct.
Distribution:
Mexico North America| Guatemala Central America|
Mexico North America| Guatemala Central America|