Calliandra Species Pages


Calliandra peninsularis


Rupert C. Barneby

13. Calliandra peninsularis Rose, Contr. U.S. Natl. Herb. 5: 135. 1897. — "[MEXICO. Baja California Sur:] ... in a garden at La Paz ... collected by Dr. Edward Palmer, January 20 to February 5, 1890 ... (n. 22)." — Holotypus, US!; clastotypus + photo, NY!. — Anneslia peninsularis Britton & Rose, N. Amer. FL 23: 58. 1928. — Calliandra sp. Rose, Contr. U.S. Natl. Herb. 1: 69. 1890, a provisional description, lacking epithet.

Anneslia lagunae Britton & Rose, N. Amer. Fl. 23: 59. 1928. — "La Laguna, Lower California, [26 Jan,] 1906, [E. W.] Nelson and [E. A.J Goldman 7455." — Holotypus, NY, on 2 sheets!; isotypus, US!.

A. brandegeei Britton & Rose, N. Amer. Fl. 23: 62. 1928. — "Sierra de Francisquito [a peak of Sa. La Victoria], Lower California, September 29,1899, T. S. Brandegee— Holotypus, Brandegee s.n., NY!. — Calliandra brandegeei Gentry, Allan Hancock Pacific Exped. 13(2): 130. 1949, quoad nom., exclus. Dawson 1165 e San José del Cabo.

Calliandra brandegeei sensu Wiggins, Fl. Baja Calif. 703. 1980.

Microphyllidious shrubs attaining ±2 m with slender terete fuscous branches, these together with lf- axes thinly (initially more densely) puberulent and often in addition granular, but glabrescent, the firm plane bicolored lfts at maturity dove-gray above, dull brown on lower face, either glabrous throughout or (sub)glabrous minutely ciliolate, the capitula arising singly or paired from axils of contemporary lvs; phyllotaxy indecisive, partly distichous, partly spiral or irregular. Stipules herbaceous, narrowly lanceolate or lance-attenuate, weakly 3-5-nerved, deciduous. Lf- formula v-vii/16-21; lf-stk of primary lvs 4—9 cm, the petiole (6-) 10-24 mm, at middle 0.5-0.8 mm diam, the ventral groove obscurely bridged between pinna- pairs, the longer interpinnal segments 4-13 mm; pinnae irregularly graduated, the rachis of longer ones 2-5 cm, the longer interfoliolar segments 1-3 mm; lft- pulvinules 0.25-0.45 x 0.2-0.3 mm, not wrinkled; lfts not or scarcely graduated, the rachis of longer ones 2-5 cm, the blades narrowly oblong or narrowly semi- ovate from obtusely semi-cordate base, deltate-apicu- late, the longer ones (3.6-)4—9 x (0.9-)1.4—3.5 mm, 2.6—4 times as long as wide; venation weakly prominulous ventrally, slenderly raised dorsally, the almost straight midrib forwardly displaced to divide blade 1:2, feebly 2-3-branched on either side from near middle or above, the one simple posterior primary nerve scarcely attaining mid-blade. Peduncles 12-23 mm, ebracteate; capitula 8—12-fld, the receptacle <1.5 mm diam, the obscurely pedicellate fls homomorphic, the bracts subulate ±1 mm or less, persistent; pedicels 0.5-0.7 x 0.3-0.4 mm; perianth 5-merous, the submembranous calyx minutely puberulent around orifice, the faintly nerved corolla thinly strigulose; calyx campanulate 1.6-2 x 1.1-1.2 mm, the tube weakly 5- nerved, the triangular teeth 0.25-0.35 mm; corolla subtubular, slightly dilated distally, 6.6-8 mm, the ovate lobes 1.4-0.9 mm; androecium 17-26-merous, 17.5-23 mm, the tube ±6 mm, the stemonozone obscure, the tassel red; intrastaminal nectary of bisexual fls 0.25 x 0.4 mm; ovary subsessile, prior to fertilization either glabrous or thinly granular. Pods 1-4 per capitulum, erect, in profile 6-11 x 0.75-1.7 cm, the sutural keels in dorsal view 1.4-2 mm wide, the stiffly papery valves reddish-brown, translucent, weakly cross-nerved, puberulent overall, low-bullate over seeds; seeds 4-8 per pod, plumply compressed- obovoid 6.5-7 x 4.7-5.2 mm, the hard testa dull tan, microscopically speckled, pleurogrammic.

On open stony hillsides and brushy banks in the oak-pine belt, 1400-1800 m, best known from Sa. La Victoria close to the Tropic line in Baja California Sur; reported by Wiggins (1980: 703) from Sa. La Giganta near 26°N; first seen by E. Palmer in cultivation as a febrifuge in the city of La Paz. — Fl. VII-I. — Tabardillo, meaning yellow fever, against which its roots provided medication.

Calliandra peninsularis is notable for indecisive phyllotaxy, elaborately decompound leaves, and relatively thin-textured pod. It differs from C. califomica, which is found in the same latitude but in desert foothills below 550 m, in more numerous pinnae and leaflets.

The glandular indumentum ascribed to C. peninsularis by Wiggins consists of small, thickened multi- cellular, so-called granular trichomes common in the genus, in this case mixed with simple hairs. The segregation of Anneslia brandegeei into a monotypic series (Britton & Rose, 1928: 51) characterized by glabrous flowers has neither fact nor reason to recommend it.

References: [Article] Barneby, Rupert C. 1998. Silk tree, guanacaste, monkey's earring: A generic system for the synandrous Mimosaceae of the Americas. Part III. Calliandra. Mem. New York Bot. Gard. 74: 1-223.