Dalea boraginea


Rupert C. Barneby

153.  Dalea boraginea Barneby

(Plate CXXXVII)

Like D. aurea in habit, 3.5-5 dm tall, the simple and monocephalous or distally few-branched stems densely pilose like the foliage with stiff, erect or subretrorse hairs up to 0.9-1.1 mm long, the foliage bicolored, the leaflets (dry) verdigris-green above, paler, more densely pilose and minutely punctate beneath; stipules subulate ± 2-2.5 mm long, becoming papery, fragile, recurved; leaves petioled, the main cauline ones 2-3.5 cm long, 3-foliolate, the broadly margined, ventrally flattened petiole and rachis together up to 9-14 mm long, the pair of leaflets obovate, the terminal one larger, broadly elliptic, 1-2.4 cm long, stalked above the pair, all prominently carinate dorsally by the pallid midrib, gland-mucronulate; spike subsessile, very dense, in fruit ±1.8 cm diam, the axis up to 4-6 cm long; bracts marcescent, narrowly lance-caudate, up to 9 mm long; calyx 6.7-7.6 mm long, stiffly setulose-barbate with spreading hairs up to 1.5-2 mm long, the tube 2.5-2.8 mm long, somewhat recessed behind the banner, the ribs prominent, the intercostal membranes charged with one (rarely interrupted) linear gland-body, the triangular-aristate, finally spreading teeth up to 4-4.9 mm long; petals yellow, incompletely known from marcescent fragments; banner 3.5-5 mm long, the slender claw 2.2-3.3 mm, the ovate-deltate blade ± 1.6-1.8 mm long, 1.8-2.2 mm wide, open ventrally at base, sometimes setulose dorsally and cili- ate, the claw sometimes bristly at base; wings 3 mm long, the claw 0.7 mm, the obliquely ovate blade 2.6 mm long, 1.7 mm wide; keel not seen; androecium (seen only faded, without anthers) 7 mm long, the filaments free for ±1.4 mm, the petal-sockets situated 4 mm above hypanthium; pod ± 3.5 mm long, quadrately obovate in profile, nearly 3 mm wide, the valves hyaline and glabrous in lower 2/3, firmer and densely pilose along the double prow; seed ± 2.5 mm long. — Collections: 2 (o).

Habitat not recorded, but to be expected on rocky limestone slopes at ± 1000 m, known only from Sierra de Santa Rosa, mpo Melchor Muzquiz, Coahuila. — Flowering in June. — Material: Coahuila. M. Muzquiz: valley of Rancho Agua Dulce, Wynd & Mueller 420 (US).

Dalea boraginea (hispid like borage) Barneby, sp. nov., D. aureae habitu similis sed caulis calycisque pube patula rigidiuscula, foliis caulinis omnibus pinnatim 3 (nec 5)- foliolatis, tuboque calycino inter costas glandula singula elongata lineari (nonnumquam hinc inde strangulata vel brevissime interrupta) obsito diversa. Petala infauste incomplete nota, sed vexilli lamina minima earn D. wrightii, foliis habituque toto absimilis, refert. — Coahuila. Melchor Muzquiz: Palm Canyon, July 9, 1936, Ernest Marsh 379. — Holotypus, GH; isotypi, F, MEXU, OKLA, TEX (2 sheets).

A strongly marked but incompletely known species, resembling D. aurea in its erect virgate stems ending in one or few massive, conelike spikes, but differing in the hispid vesture of stem and calyx, in the pinnately trifoliolate leaves, and in the single linear gland on each intercostal membrane of the calyx. Its close relationship in sect. Cylipogon will require reassessment when the epistemonous petals become better known. Marsh collected D. aurea also, several times, in Muzquiz municipality (Nos. 56, GH, TEX; 1281, F, GH, TEX; 2145, GH, OKLA), but these specimens are characteristic of their species and show no hint of passage toward D. boraginea. The small vexillum suggests that of D. wrightii, a smaller plant with always more numerous leaflets. The only trifoliolate member of sect. Cylipogon known to occur in the same region is the dwarf C. laniceps, including stem and spike less than 1 dm tall.

References: [Article] Barneby, Rupert C. 1977. Daleae Imagines, an illustrated revision of Errazurizia Philippi, Psorothamnus Rydberg, Marine Liebmann, and Dalea Lucanus emen. Barneby, including all species of Leguminosae tribe Amorpheae Borissova ever referred to Dalea. Mem. New York Bot. Gard. 27: 1-892.

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