Dalea chrysophylla


Rupert C. Barneby

94.  Dalea chrysophylla Barneby

(Plate LXXXIV)

Suffruticose, the slender, sparsely branching, remotely gland-verruculose stems apparently ascending, up to 3-4 dm tall, except for old wood densely permanently pilose-tomentulose throughout with fine ascending and spreading ± tangled hairs up to ± 0.7 mm long, the herbage probably silvery when fresh, golden when dry, the leaflets equally pubescent both sides, punctate under the vesture beneath; leaf- spurs 0.4-0.7 mm long; stipules linear-setiform but soft, 3-5 mm long; intra- and post- petiolular glands very small, concealed by vesture; leaves shortly petioled, the main cauline ones 2-3 cm long, with very narrowly margined rachis and 4-5 pairs of sub- sessile, oblong-elliptic or -obovate, apically penicillate and gland-mucronate, shallowly concave, dorsally keeled, thick-textured leaflets 4-7 mm long, the terminal one usually stalked; peduncles, except the first of each primary axis, terminal to lateral branchlets, 2-6 cm long; spikes moderately dense, the flowers ± 3-ranked after anthesis, ovoid becoming cylindroid, without petals 8 mm diam, the densely villosulous axis becoming 1.5-5 cm long; bracts persistent, 4-4.5 mm long, densely pilosulous dorsally, the navicular body 2-2.7 mm long, in profile 0.9-1.1 mm wide, contracted into a subulate tail ± 1.5-2 mm long; calyx 4.4-4.5 mm long, pilosulous from base upward, the tube (measured to dorsal sinus) 2.8-3 mm long, recessed behind banner, the orifice oblique, the livid ribs filiform, the membranous flat intervals charged with 5-9 tiny glands arranged in 2-3 rows, the teeth deltate-apiculate, the dorsal one longest, 1.5-1.8 mm long (1.2-1.5 mm shorter than tube), the ventral pair shorter and broader; petals bicolored, the whitish, gland-sprinkled and -tipped banner early rubescent, the epistemonous ones bright blue, perched low on androecium (1.4-2 mm above hypanthium), the keel gland-tipped; banner 4.6-4.8 mm long, the claw 2 mm, the broadly ovate-cordate hooded blade 2.9 mm long, 3.6 mm wide; wings 4.1-5 mm long, the claw 1.1-1.4 mm, the obliquely ovate blade 3.1-4 mm long, 1.6-2 mm wide; keel 5.2-5.6 mm long, the claws 1.6-1.8 mm, the blades 3.7-4 mm long, 2.2-16 mm wide; androecium 10-merous, 5.5-6 mm long, the longer filaments free for 1.6-2 mm, purple distally, the connective gland-tipped, the anthers 0.3-0.45 mm long; pod 2.4 mm long, in profile half-obovate, the style-base terminal, the valves hyaline and glabrate in lower half, thence very thinly papery, gland-sprinkled, pilosulous; seed ±1.7 mm long. — Collections: 2 (o).

Habitat little known, sometimes in old fields, near 2100 m, known only from the w. slope of the Andes in Azuay, Ecuador. — Flowering September to November.—Material: ECUADOR.: W. Lobb, s.n. (K, herb. Hook., reed. 1867). Azuay: typus.

Dalea chrysophylla (golden-leaved, of the dense pubescence) Barneby, sp. nov., inter congeneres andinas parvifloras pube omnium partium piloso-tomentosa valde densa in sicco aurata praestans, a sympatrica D. ayavacensi var. killipii subsimiliter tomentosa sta- tura minori ac calyce brevidentato ab ipso basi pilosulo, utroque tubi intervallo glandulis numerosis parvulis sparsis nec in 2 (3) lineares confluentibus obsito facilius separata.— Suffrutices 3-4 dm altae undique densissime piloso-tomentosae, pube exsiccata aurea; folia majora 2-3 cm longa, foliolis 4-5-jugis oblongo-obovatis 4-7 mm longis; spicae densiusculae anguste ovoideae demum cylindricae, petalis ablatis 8 mm diam, 1.5-5 cm longae; bracteae persistentes 4-4.5 mm longae medium versus in caudam brevem con- tractae; calycis tubus ± 2.9 mm longus ad orem obliquus, dentes deltato-apiculati 1.5-1.8 mm longi; petala bicoloria, vexilli ± 4.7 mm longi lamina albida mox rubescenti, episte- monea coerulea carinae laminis 3.7-4 mm tantum longis. — ECUADOR. Azuay: old field, 90 mi s. of Cuenca, 7000 ft, oct 2, 1944, Ira L. Wiggins 10,838. — Holotypus, NY; isotypus, US.

This suggests a densely pannose-tomentulose version of D. humifusa or D. exilis. The only sympatric dalea similarly pubescent is D. killipii var. killipii, very like it in the individual leaf but at the same time very different in its greater stature, long calyx-teeth, and externally glabrous calyx-tube charged in each interval with a single linear gland. The type-collection of D. chrysophylla was distributed as D. sericophylla Ulbr., to which it would key following Macbride’s revision of the Andean species, but differs in its shorttailed bracts and smaller flowers. The poorly known D. sericophylla has been collected only in northern Peru (Cajamarca) and is interpreted herein as a variety of the middle- Peruvian D. weberbaueri. A third densely pubescent dalea was collected long ago in the province of Chachapoyas, but in specimens (Mathews in 1835, OXF, K) inadequate for thorough study. It appears to differ from D. chrysophylla in its substantially larger corolla, but the value of these differences will obviously need reevaluation as more material becomes available.

References: [Article] Barneby, Rupert C. 1977. Daleae Imagines, an illustrated revision of Errazurizia Philippi, Psorothamnus Rydberg, Marine Liebmann, and Dalea Lucanus emen. Barneby, including all species of Leguminosae tribe Amorpheae Borissova ever referred to Dalea. Mem. New York Bot. Gard. 27: 1-892.

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