Dalea cylindrica var. nova


Rupert C. Barneby

80d.  Dalea cylindrica Hooker var. nova (Ulbrich) Barneby

(Plate LXXXII)

Herbaceous becoming softly fruticose, highly variable in stature, erect, diffuse, or in exposed sites even humifuse, at anthesis (0.3) 0.5-2 m tall, the stems at least distally verruculose; primary leaves (1.5) 2-5.5 cm long, the 6-10 (11) pairs of leaflets 5-13 (17) mm long, glabrous above; calyx-tube thinly pilose with lustrous hairs up to 0.8-1.5 mm long, the ribs early becoming stout and prominent, the intervals charged with 3-5 small, yellow or golden glands, the apically glabrate teeth stiff and corneous; banner whitish (rubescent), either glabrous or puberulent dorsally; epistemonous petals usually vivid cobalt blue, sometimes violaceous; spikes (1.5) 2-9 cm long.—Collections: 34 (o).

Dry stony crests and hilltops, hill pastures, and banks of eroded gullies, descending into scrub and valley flats, in exposed sites diffuse or prostrate becoming erect and bushy in protected ones, mostly 2150-3000 m, but in Ecuador descending in dry barrancas to 1200 m, locally abundant in the inter-Andean valleys of centr. and n. Peru, from Junin n. to Amazonas and Piura, and n. through Ecuador as far as Chimborazo, from La Libertad n.-ward extending across the Divide to the edge of the Pacific slope, s.-ward entirely e. of the Divide. — Flowering nearly all year — Representative: ECUADOR. Chimborazo: A. S. Hitchcock 20,745 (NY, US). Canar: Prieto P-174 (K, NY). Azuay: Camp 2528 (F, NY, US); Rose, Pachano & Rose 22,815 (NY, US). Loja: Seemann 753 (K); D. H. Knight 755 (NY, WIS). PERU. Piura. Huancabamba: Sandeman 4272 (K, OXF). Chachapoyas. Chachapoyas: Wurdack 415 (NY, UC, US); Mathews s. n. (F, K, NY). Cajamarca. Cutervo: Sandeman 4075 (K, OXF); Ferreyra & Acleto 15,351 (NY, US). Chota: Stork & Horton 10,053 (F, K, UC). Celendin: Ferreyra 15,046 (NY, US); Hutchinson & Wright 5126, 5262 (NY). Cajamarca: Ferreyra 3229, 3284 (US); Hutchinson & Wright 5057 (NY, UC, US). La Libertad. Otuzco: Lopez Miranda 0608 (US). Huanuco. Ambo: Mathews 930 (K, OXF); Sawada P.112 (F). Huanuco: Ferreyra 6694 (US). Junin: Uspachaca, Macbride 1302 (F).

Dalea cylindrica Hook. var. nova (Ulbr.) Barneby, stat. nov., based on D. nova (new) Ulbr., Feddes Repert. 2: 5. 1906. — "Peruvia: Depart, et provincia Cajamarca, supra San Pablo 2400- 2700 m. (A. Weberbauer, Fl. v. Peru no. 3838 —spec. fl. et fr. coll. 26. april 1904.)." — Holotypus, formerly B, survives as Field Neg. 732, F, NY, US! — Parosela nova (Ulbr.) Macbr., Field Mus., Bot. 4: 110. 1927.

Parosela sawadae (Mosaho Sawada, collected in Peru ca 1927) Macbr., Field Mus., Bot. 11: 24. 1931. — "Peru: Huanuco, Sawada P 11...San Rafael, Dept, of Huanuco, Sawada P 112." —Holotypus, Sawada P-119 (not P-11), F! paratypus (Sawada P 112), F! — Dalea sawadae (Macbr.) Macbr., Candollea 7: 223. 1937.

The material assembled under the heading of var. nova is diverse in superficial appearance. An observant collector (F. Woytkowski 5, F) records that the species is common around Celendin in Cajamarca, where it occurs both in the valley and on the higher hills. In hedges where protected from wind and grazing, the plants shoot up to a height of nearly 14 dm, but in open hill country they assume a diffuse or even prostrate form, running together in "wide clusters like heather," seldom over 5 dm tall. Primary leaves on long branches of the taller type are composed of up to 9-11 pairs of realtively large leaflets; and the first spike of each primary axis is usually leaf-opposed and long-pedunculate. In the lowly phase the majority of inflorescences are borne terminal to short branchlets set with shorter leaves composed of only 3-7 pairs of leaflets. In addition to these seasonal and edaphic variants there is an expectable instability in vesture and in density of glands on the bracts, peduncles and young stems. As a result it is not easy to characterize var. nova in exact terms. Over much of its range in Peru it coexists with var. haenkeana, which resembles its dwarf aspect, but differs in having the calyx nearly of var. cylindrica; and with the superficially rather similar D. coerulea var. longispicata. In Ecuador it enters the range of D. coerulea var. coerulea. The two entities last mentioned have often been confused with var. nova, but can be separated as follow:

1.  Calyx-teeth pilose-plumose from base to apex, the tips surpassed by at least some hairs; dorsal calyx-tooth 2.7-5.5 mm long, at least as long and commonly much longer than the tube, flexuous in age, the marginal nerves not greatly thickened. D. coerulea.

1.  Calyx-teeth pilose at base and internally, the glabrous tips surpassing the longer hairs, the haifs ordinarily reduced in length upward; dorsal calyx-tooth 1:7-2.7 (3) mm long, at least 0.2 mm shorter than the tube, erect and stiffened in age by the corneous marginal and costal nerves. D. cylindrica var. nova.

References: [Article] Barneby, Rupert C. 1977. Daleae Imagines, an illustrated revision of Errazurizia Philippi, Psorothamnus Rydberg, Marine Liebmann, and Dalea Lucanus emen. Barneby, including all species of Leguminosae tribe Amorpheae Borissova ever referred to Dalea. Mem. New York Bot. Gard. 27: 1-892.

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