Dalea quercetorum


Rupert C. Barneby

74.  Dalea quercetorum Standley & L. Williams

(Plate LXXVI)

Coarse, softly woody herbs and shrubs variable in habit and stature, often flowering the first year as monopodial herbs 0.6-1.5 m tall, later several stemmed from a caudex, or ultimately true shrubs up to 4.5 m tall, the foliage and the young, livid or greenish, inconspicuously verruculose stems densely silky-pilose or -pilosulous with fine flexuous hairs up to 0.5-1.5 mm long, the leaflets equally pubescent both sides, densely gland-sprinkled beneath, the spiciform racemes mostly terminal to leafy branchlets, forming an open corymbose panicle; leaf-spurs 0.7-2.5 mm long; stipules linear-caudate, pale or livid, becoming stiff and fragile, 3.5-7 mm long, densely pilosulous and sparsely verruculose dorsally; intrapetiolular glands 2-4, spiculiform; post- petiolular glands small but prominent, blunt; leaves shortly petioled, the main cauline ones (2.5) 3-9 cm long, with stiff, narrowly margined rachis and 3-7 pairs of oblong: or ovate-elliptic to broadly oblanceolate, abruptly short-acuminate, flat, dorsally carinate leaflets up to 1-2.5 (mostly 1-1.5) cm long; spikes (racemes) moderately dense, many-flowered, becoming oblong-cylindroid, without petals 12-15 mm diam, the subcontiguous flowers widely spreading-ascending, the densely pilosulous axis (1) 2-14 cm long; bracts deciduous, 3-6(8) mm long, the body broadly lanceolate, navicular, keeled dorsally, castaneous except for scarious-margined base, 2-4 (4.5) mm long and in profile 1-1.6 mm wide, tapering or abruptly contracted into a livid tail 1.5 2.5 (3.5) mm long, the whole pilosulous dorsally throughout or almost so, glandless or minutely gland-sprinkled, glabrous within; pedicels up to 0.3-0.7 mm long, sometimes almost 0; calyx 5.5-7.5 mm long, finely and densely pilose with spiral hairs up to (0.8) 1-1.4 mm long, the tube 2.7-3.7 mm long, somewhat recessed behind banner, the orifice moderately oblique, the slender, livid-castaneous or rarely pallid ribs becoming prominent, the membranous flat intervals charged with many small scattered, golden or honey-colored glands, the lance- or triangular-cuspidate gland-tipped and -spurred teeth unequal, the dorsal one longest, 2.2-4 mm long (as long to 1.2 mm shorter than tube), the ventral pair shortest; petals bicolored, the banner white distally, rubescent, its basal lobes blue-tipped, charged with a few glands in the eye and dorsally above top of claw with a small tuft of hairs, sometimes like the keel minutely gland-tipped, the epistemonous petals deep vivid blue fading violet-blue, paler along outer edges, perched well below middle of androecium (± 2.2-3.8 mm above hypanthium); banner 6-7 mm long, the claw 3.2-3.8 mm, the suborbicular emarginate blade callous but not recessed into a comet at base, 3-3.8 mm long, 3-4 mm wide, its lobes inflexed and adherent to form lateral pockets; wings 5.5-5.8 mm long, the claw 1.2-1.7 mm, the ovate-oblong blade 4.3-4.7 mm long, 2-2.2 mm wide; keel 6-7.3 mm long, the claws 1.7-2.4 mm, the elliptic blades 4.7-5.5 mm long, 2.7-3.1 mm wide; androecium 10-merous, 7-9 mm long, the longest filaments free for 1.7-2.5 mm, the connective gland-tipped, the anthers 0.55-0.7 mm long; pod in profile triangular or triangular-obovate, 2.6-3.4 mm long, the style-base latero-terminal, the prow slenderly keeled, the valves hyaline in lower 2/3, thence thinly papery, minutely gland-sprinkled, pilosulous with fine spiral hairs; seed castaneous, 2-2.9 mm long.—Collections: 10 (o).

Habitat little known, but at least sometimes in oak-pine woods, ± 1500-1700 m, apparently local and discontinuous: highlands of Chiapas, Mexico (mpos Amatenango del Valle, Comitan, La Trinitaria); s.-e. Guatemala, mostly within or near the drainage of Rio Motagua (dptos Baja Verapaz, Jalapa, Zacapa), and Honduras (dptos Comayagua, El Paraiso, Intibuca).— Flowering November through February.— Material: MEXICO. Chiapas: Miranda 5014 (MEXU); Matuda 6022 (F, MEXU); Breedlove & Raven 8363 (US). GUATEMALA. Baja Verapaz: Molina & Molina 12,353 (F, NY). Jalapa: Steyermark 33,127 (F). Zacapa: Steyermark 42,936 (F), 43,186 (F, NY). HONDURAS. Comayagua: Yuncker et al 6238 (F, sterile); El Paraiso: Molina 13,075 (F, NY). Intibuca: Molina & Molina 24,287 (NY); Standley 25,191, 25,268 (F, topotypi).

Dalea quercetorum (of oak-woods) Standl. & L. Wms., Ceiba 1: 82. 1950.— HONDURAS: Dept. Intibuca: Cerca de La Esperanza...En. 31-Feb. 12, 1950, Standley 25454..."—Holotypus, olim in herb. Esc. Agr. Panam., hodie US!

Dalea leucosericea sensu Standi. & Steyerm., Field Mus. Bot. Ser. 24s: 213. 1946; non Parosela leucosericea Rydb.

A coarsely leafy dalea, readily recognized by floral characters: glands in intercostal membranes of calyx numerous, scattered; petals except for the partly white banner vivid blue; banner about as long as keel and bearing on back a small tuft of hairs. Of sympatric species only D. dispar has a similar banner, but combined with greenish-yellow petals and vertically uniseriate glands in the calyx-membranes. Both these species flower precociously as tall, simple-stemmed herbs branched distally into a panicle of spikes but later become soft shrubs, D. quercetorum reportedly up to 4.5 m tall. The phases of maturity are well shown by Steyermark 42,936 (monopodial herb), Steyermark 33,127 (suffruticose, with several virgate stems), and several collections consisting of branches taken from shrubs of variable stature. Independently of stature, the pubescence of the stems varies from relatively thin and forwardly subappressed, not fully concealing the chestnut- purple epidermis, to very dense, horizontally spreading, and shaggy. The pedicels, ordinarily well-developed, are sometimes short or obscure. The leaflets of some pressed specimens show a characteristic sleep-movement, becoming reflexed, the members of each pair back to back, and retrorsely imbricated.

As implied in the synonymy, I have referred to D. quercetorum, described from Honduras, the Guatemalan plants assigned by Standley & Steyermark to D. leucosericea, a species sometimes similar in its dense, when fresh silvery (in dried specimens early tarnished) pubescence of long, glossily silky hairs, but different in the always sessile calyces subtended by flatter, internally silky bracts, uniseriate glands in the calyx-intervals, pinkish-lavender petals, and dorsally glabrous banner substantially shorter than the detached keel. The known ranges of D. leucosericea, not known to occur southeast of central Oaxaca, and D. quercetorum, with its western limit in highlands of eastern Chiapas, are so far apart that no further confusion of the two seems likely,

References: [Article] Barneby, Rupert C. 1977. Daleae Imagines, an illustrated revision of Errazurizia Philippi, Psorothamnus Rydberg, Marine Liebmann, and Dalea Lucanus emen. Barneby, including all species of Leguminosae tribe Amorpheae Borissova ever referred to Dalea. Mem. New York Bot. Gard. 27: 1-892.

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