Marina sarodes


Rupert C. Barneby

30.  Marina sarodes Barneby

(Plate XXI)

Tall, slender, erect annual herbs from an orange root, paniculately branching distally (with habit of M. scopa), glabrous throughout, the almost eglandular, smooth stem livid becoming stramineous distally, the foliage bicolored, the leaflets green above, pallid and sparsely punctate beneath, lineolate; leaf-spurs 0.4-0.7 mm long; stipules 1-1.5 mm long, gland-fimbriolate; intrapetiolular gland small; post-petiolular

glands small but prominent, conic, orange; leaves subsessile, the main cauline ones 2.5-4 cm long, with narrowly margined rachis and 8-16 pairs of narrowly oblong or oblong-oblanceolate, flat, minutely gland-crenulate leaflets 3-6 mm long, the leaves of the inflorescence abruptly smaller, 5-15 mm long, with only 1-3 (4) pairs of oblanceolate to narrowly oblong leaflets; racemes 5-11-flowered, the lower flowers separated by a distance longer than their pedicels, the distal ones more crowded, the axis becoming 1-2.5 cm long; pedicels 0.6-1 mm long; calyx 3.4-3.6 mm long, the firm, purplish tube 1.9-2.1 mm, charged in each interval with a row of 2-3 small glands, the teeth green with narrow, pallid, erose margins, dissimilar, the dorsal one longest, narrowly ovate, 1.4-1.6 mm long, the ventral pair shorter, broadly obovate; petals bicolored, the banner white with purple-tipped basal lobes, rubescent, the inner bright magenta-purple, perched 1.5-2 mm above hypanthium rim, the blades of banner and keel sparsely gland-sprinkled; banner 3.2-3.7 mm long, the rather thick claw 1.3-1.6 mm, the obtusely deltate-obcordate blade recurved through ± 50°, closed across the top of the claw to form a shallow comet; wings 4.7 mm long, the claw 0.3 mm, the blade horizontal to keel at full anthesis, broadly obovate, 4.6 mm long, 3 mm wide; keel 6 mm long, the claw 1 mm, the broadly oblong-obovate blade 5.3 mm long, 3.1 mm wide; androecium 10-merous, 6 mm long, the longest filament free for ± 1.5 mm, the connective gland-tipped, the anthers 0.6 mm long; ovary charged with small glands arranged in 1 transverse and 1 subvertical crescent, the forming pod compressed, not seen ripe. — Collection: 1 (typus).

Brushy slopes in oak woodland, ± 900 m (3000 ft), known only from one station in the e. end of the Transverse Volcanic Belt in s. Nayarit. — Flowering October and November, possibly later.

Marina sarodes (broomlike, in habit resembling the broom marina M. scopa) Barneby, sp. nov., inter ser. Crenulatas radice annua, caule solitario erecto ultra medium panicu- latim ramoso insignis. Annuas sect. Carroae species M. minutifloram necnon M. scopam habitu simulans sed ab ilia caulibus elatis floribusque majoribus, ab hac vexilli lamina ampliori emarginata peltata, ab ambabus leguminis ut videtur compressi valvulis more seriei suae glandulis lunatim ordinatis nec conspersis ornatis gravius discrepat. — NAYARIT. Jalisco: steep brushy oakwoods, 3000 ft., 11 mi. s. of Jalisco, oct 8, 1965, Ripley & Barneby 14,044. — Holotypus, NY; isotypus, CAS.

Granted that M. sarodes is correctly referred to ser. Crenulatae, as suggested by the crescentic gland-patterns on the young pod, it is a most distinct species, easily and immediately distinguished from M. crenulata and M. dispansa by its annual, monopodial growth- habit, and from M. neglecta further by its many pairs of leaflets in main cauline leaves. In habit it resembles some species of sect. Carroa, notably M. scopa and M. minutiflora, but differs from the latter in its taller stems and much larger flower, and from M. scopa, often similar in stature, in the peltate blade of the much larger banner. Until the truly ripe fruit of M. sarodes is discovered, the affinities of the species will remain open to controversy, but its status as a species seems secure. All annual members of ser. Unifoliolatae differ from M. sarodes in the pectinate ciliolation of the calyx-ribs and a banner-blade open at the base.

Only a few plants of M. sarodes were seen at the type-locality, growing on the precipitous bank of a ravine, among blocks of volcanic rock in light shade of oaks, associated with Calochortus hartwegii Bth. and the rare annual Dalea dipsacea. At the time my attention was distracted by discovery of the latter, and I mistook M. sarodes for the common and weedy M. scopa. Possibly other collectors may have passed it by for the same reason.

References: [Article] Barneby, Rupert C. 1977. Daleae Imagines, an illustrated revision of Errazurizia Philippi, Psorothamnus Rydberg, Marine Liebmann, and Dalea Lucanus emen. Barneby, including all species of Leguminosae tribe Amorpheae Borissova ever referred to Dalea. Mem. New York Bot. Gard. 27: 1-892.

Multimedia: