Heuchera hapemanii Coulter & Fisher, Bot. Gaz. 17: 348. 1892. Sullivantia hapemanii (Coulter & Fisher) Coulter, Bot. Gaz. 17: 421. 1892. Sullivantia oregana var. hapemanii (Coulter & Fisher) C. 0. Rosend., Bot. Jahrb. 37, Biebl. 83: 60. 1905. TYPE: U.S.A. WYOMING: "Big Horn Mountains," Hapeman s.n. (HOLOTYPE: GH!; IsoTYPEs: F!, US!).
Sullivantia halmicola A. Nelson ex Small, N. Amer. Fl. 22: 122. 1905. TYPE: U.S.A. WYOMING: "Hat Six Creek, Casper Mountain," 7 Aug 1898, E. Nelson 5032 (HOLOTYPE: NY!; ISOTYPES: ILL!, MO!, RM!, US!).
Moist cliffs, in close proximity to waterfalls, and along streams, typically occurring on limestone. Big Horn Mountains and Laramie Mountains (the latter represented by a single population from Casper Mountain), Wyoming and Montana. Flowering occ,urs from June through
In 1905 Small described a new species, S. halmicola, attributed to A. Nelson and based on E. Nelson's collection from Casper Mountain, Wyoming. This species was distinguished from S. hapemanii chiefly on the basis of size differences. That this was a dubious distinction is illustrated by the fact that floras published soon after the publication of S. halmicola reduced this species to synonymy with S. hapemanii (Coulter & Nelson, 1909; Britton & Brown, 1913). Rosendahl (1927), however, retained S. halmicola as a distinct species. This study has shown that there is no persuasive evidence from morphology, crossability, or flavonoid chemistry to suggest that these two species should be retained. Sullivantia halmicola has, therefore, been reduced to synonymy.
The precise year of collection of the type of S. hapemanii is not given on the label.
According to the protologue for S. halmicola, the type was collected at "Hat Six Creek, Casper Mountain, Wyoming, Aug. 7, 1898, by E. Nelson, no. 5032," but no holotype was designated. However, the fact that only the type collection of S. halmicola deposited at NY is stamped "examined for North American Flora" effectively designates this specimen as the holotype
Specimens examined: U.S.A. Montana. Carbon Co.: Dorn 2674 (RM); Soltis & Hammond-Soltis 1033 (IND, MONTU). Wyoming. Big Horn Co.: Fassett s.n., in 1939 (WIS); Ownbey & Ownbey 1875 (COLO, GH, ISC, MIN, MO, RM, UC, UMO, WS); Soltis & Hammond-Soltis 1029 (IND, RM). Johnson Co.: Soltis & Hammond-Soltis 1024 (IND, RM); Weber et al. sn., in 1974 (COLO). Natrona Co.: Nelson 5032 (ILL, MO, NY, RM, US). Sheridan Co.: Garber 10771 (GH, MIN, MO, RM, UC); Hayden s.n., in 1859 (MO); Hapeman 907 (GH); Hapeman s.n., in 1892 (F, US); Hapeman s.n., in 1896 (NY, WIS); Hapeman s.n., in 1906 (BUT, F, GH, ISC, MIN, MU, NY, RM, UC, UMO, WS, WTU); Hapeman s.n., in 1909 (IND); Nelson 2302 (GH, MO, NY, US); Soltis & Hammond-,Soltis 1027, 1028 (both IND, RM); Willets 261 (RM).