Monographs Details: Randia aculeata var. dasyphylla Steyerm.
Authority: Maguire, Bassett. 1972. The botany of the Guayana Highland--part IX. Mem. New York Bot. Gard. 23: 1-832.
Family:Rubiaceae
Description:Latin Diagnosis - A var aculeata atque var dasyclada foliorum paginis subtus manifeste pilosulis necnon costa media nervis lateralibusque pilis patentibus abundanter munitis recedit; hypanthio atque calycis tubo plerumque copiose pubescenti.

Distribution and Ecology - Distribution. Venezuela, Colombia, and Trinidad. TRINIDAD. Moruga, 19 Mar 1921, Britton & Broadway 2459. VENEZUELA. Bolivar: between Hato Corumo and Las Chicharras, NE of Tumeremo, 1 Jun 1960, Steyermark 86242; Alto playa de la sabana region, Upata-El Palmar, 11 Dec 1959, Ramia 2087; sabanas entre Guasipati y El Manteco, 21 Feb 1960, Ramia 2267. Nueva Esparta: El Valle, Margarita Island, 31 Aug 1903, Johnston 280; Cerro Matasiete, Margarita, 20 Aug 1953, Foldats 2719. Miranda: Rio Chico, 1 Jun 1923, Jahn 1258. Carabobo: road from Puerto Cabello to San Felipe, Guaremales, alt 10-100 m, 29 May 1920, Pittier 8849. Yaracuy: La Mingoya, San Felipe, 26 Oct 1949, Trujillo & Fernandez 501; alrededores de Corumbo, via Nirgua, alt 750 m, 19 Apr 1968, Rojas 192. COLOMBIA. Magdalena: costa del Caribe, valle del Río Cesare (parte occidental), al oeste y sur de Los Venados, 10° lat N, 73° 42' W, alt 60 m, 11-13 Feb 1961, Dugand 5583.

Discussion:

Type. Savanna-chaparral, between Hato Corumo and Las Chicharras, NE of Tumeremo, BolĂ­var, Venezuela, 1 Jun 1960, Julian A. Steyermark 86241 (holotype VEN).

This variety is distinguished by the conspicuously pilosulous lower surface of the leaf blades as well as by the abundant spreading pubescence on the midrib and lateral nerves of the lower side. Moreover, the hypanthium and calyx tube are generally more abundantly pubescent, as is also usually the fruit, although it may lose hairs in age, becoming glabrate as in Britton & Broadway 2459. In a mature state the pubescence is well shown in Trujillo & Fernandez 501. The leaves have the general shape of those of R. aculeata var aculeata, ranging from obovate to narrowly elliptic to oblanceolate, and from rounded to subacute at apex. They may be broadest at the middle or in the upper third, and are usually attenuate or contracted to the base into a decurrent subpetiolate portion 1-11 mm long. They vary from 10-50 X 6-22 mm.