One species, Gomidesia bonnettiasylvestris Steyermark, has been described from
the Guayana Highland area. It is treated and briefly described above under Myrcia
sect Myrcia. The genus Gomidesia is a group of about 40 or more species, mostly
natives of the south-Brazihan region. Legrand has recently published a synopsis of
practically the entire genus (Las especies tropicales del genero Gomidesia. Com. Bot.
Mus. Hist. Nat. Montevideo 3(37): 1-30. 1959 ["31 Dec 1958"]).
The diagnostic character of Gomidesia is wholly in the anthers; whereas in Mvrcia
the anthers are bilocular, longitudinally dehiscent and explanate, in Gomidesia they
are incompletely (or in one species wholly) 4-locular; the interior sac of each theca
is somewhat displaced distad, and seems to open extrorsely at the tip, and the exterior
sac seems to open introrsely at the base. The differentiation in the anthers may be
barely perceptible (i.e., in species otherwise referable to Myrcia), or it may be considerable.
In the most extreme situation the anthers have four apical pores (the
genus Cerquieria). Bentham summarily reduced Gomidesia and Cerquicria to Mvrcia,
on the ground that the differences were "so difflcult to appreciate, and so little in
accordance with habit." Legrand's work indicates, however, that Gomidesia occupies a natural range, mostly in southern Brazil with a few species ranging as far north as
Bahia and one only to the West Indies. Although the anther-character is the only
thoroughly consistent one distinguishing the genus from Myrcia, most of the species
are recognizable by the tawny or reddish-brown silky, somewhat appressed pubescence
that covers the inflorescence and the flowers.