Monographs Details: Glossarion
Authority: Carlquist, Sherwin. 1957. Anatomy of Guayana Mutisieae. Mem. New York Bot. Gard. 9 (3): 441-476.
Discussion:Flowers of the monotypic genus Glossarion (fig. 116) are of particular interest in that they are ligulate in form, all the lobes being united except at the tips (fig. 116 A-C). Although the lobes are histologically united, the vascular anatomy suggests that the corolla form is basically actinomorphic, with some degree of modification. As the corolla in figure 116 A shows, some of the lateral veins unite beneath the short sinuses at the "ligule" tip. It seems significant, however, that each of the five veins of the corolla-tube divides into two laterals at the same level as the vein beneath the single deep sinus in the corolla. Portions of two other corollas are shown (fig. 116 B, C) to indicate the variation in venation at the corolla tip. In each of these, however, ten laterals are present in the more basal portion. Occasionally one of the outer lobes may be separate for a short distance (fig. 116 B). The vascular pattern of the corolla, then, shows little modification from an actinomorphic form, although the tissues between these lobes are continuous. This is a stage in union of corolla lobes beyond that found in Fitchia (Carlquist 1957 b), where sinuses are Justologically formed between the lobes, although only the adaxial sinus usually opens during anthesis. Stamen traces unite with their associated corolla bundles in the achene top. In addition to the five exterior bundles, which continue into the corolla, five additional bundles, terminating at the achene summit, form the complement of ten exterior achene bundles (fig. 116 E). The style of Glossarion (fig. 116 D) contains four bundles, uniting to two above. These four bundles continue downward into the achene as interior bundles. Occasionally one of the dorsiventral interior bundles may unite with a nearby exterior bundle (fig. 116 H). The lateral pair, however, follow the bands of stigmatoid tissue to the base of the achene, uniting with the ovule trace (fig. 116 I). A single vascular strand results from the union of all of the bundles at the achene base. The ovule trace is simple.