Monographs Details: Coussarea longiflora var. benensis (Britton ex Standl.) Steyerm.
Authority: Maguire, Bassett. 1967. The botany of the Guayana Highland--Part VII. Mem. New York Bot. Gard. 17: 1-439.
Family:Rubiaceae
Description:Distribution and Ecology - Amazonian Brazil and Bolivia. BRAZIL. Acre: Basin of Rio Purus, near mouth of Rio Macauhan (tributary of Rio Yaco), lat. 9°20'S, long. 69°W, Krukoff 5407, 5544. BOLIVIA, junction of rivers Beni and Madre de Dios, Rusby 2402 (type of Coussarea benensis).

Discussion:

Coussarea benensis Britton ex Standley, Field Mus. Publ. Bot. 7: 296. 1931.

Type. Junction of rivers Beni and Madre de Dios, Bolivia, Aug 1886, H. H. Rusby 2402 (type of Coussarea benensis).

There is some intergradation between the two varieties treated here, but they appear to be separable for the most part. In the type material of Coussarea benensis there is a definite, usually well-developed peduncle and pedicellate flowers with pedicels 1.5-2 mm long and thick, and the peduncle 5-10 mm long, whereas in typical C. longiflora the flowers are sessile and lack a peduncle, if one may judge by the photograph of the type. This photo shows subulate, elongate calyx- lobes on the central flowers, and the calyx-lobes are longer and narrower than in the type collection of C. benensis. In C. benensis the type collection shows the elongated calyx-lobes of the central flowers only 2-4.5 mm long by 0.5-0.6 mm broad, and varying from ligulate-linear to ovate-oblong with obtuse to subobtuse tips. However, some of the calyces within a group in C. benensis are without the elongated calyx-lobes or more than one flower within a group may present elonĀ­gated lobes. Standley indicated that C. benensis is synonymous with C. brevicaulis Krause, but, as shown in the present treatment, it is related to, if not entirely conspecific with C. longiflora. In C. brevicaulis the calyx and hypanthium are much shorter and the flowers are more numerous and fasciculately clustered than in C. longiflora and C. benensis. Also, the calyx-teeth of C. brevicaulis are all very short instead of well developed in the central flowers as in C. longiflora and C. benensis.

Distribution:Brazil South America| Bolivia South America|