Bellardia tontanea (H.B.K.) Roem. & Schult., Syst. Veg. 3: 205. 1818.
Bellardia repens Willd., Sp. Pl. 1: 626. 1797.
Type. “Prope Buenavista, Caracas et Popayan, alt 500-900 hex.” [Venezuela and Colombia] Humboldt cfc Bonpland.
I am assigning this name to those plants with subappressed incurved cauline hairs, hirtellous exterior of corolla-lobes, hypanthium with dense pubescence of incurved-ascending hairs, and lower surface of leaf blades rather moderately pilose with short hairs.
The type locality for this species is not clear, since Buena Vista and Popayan refer to Colombian localities, while Caracas refers to Venezuela. The only Coccocypselum collections from the Coastal Cordillera of Venezuela involve three taxa: (1) C. lanceolatum (R. & P.) Pers., (2) plants with spreading- pubescent stems and loosely villous-pubescent leaf-blades which correspond to “Coccocypselum hirsutum Bartl. ex DC.” of Trinidad origin, and (3) plants with subappressed-pubescent stems and leaves corresponding to a collection of Jenman 4547 from British Guiana identified at Kew Herbarium as C. tontanea. It is to the third or last category for which I have adopted the name C. tontanea. Collections with more densely pubescent leaves and with more elevated nerves beneath occur in the Gran Sabana of Venezuela (Steyermark 59411) and in Colombia near the type locality or cited locality of both C. lanceolatum and C. tontanea at “Buena Vista,” where collected by Pennell (no. 1676). These more densely pubescent-leaved specimens appear to belong to an undescribed species, which is indicated below.
TRINIDAD. Mt. Tucutche, Friend 15. BRITISH GUIANA. Butukari, Gleason 719; Rockstone, Gleason 542, 575; Koreai Creek, Linder 155; Huradaiah, Moruka River, Pomeroon District, De La Cruz 1023; Pomeroon River, Pomeroon District, De La Cruz 3200; Waini River, Northwest District, lat 8°20'N, long 59°40'AAT, De La Cruz 3755; Macaseema path to Cabacaburi, Beeket s.n., Iramai-pang, Forest Dept. Record No. WB 593; upper Mazaruni River basin, Kamarang River, base of Utschi Falls, ca 640 in, S. S. & C. L. Tillett & R. Boyan 45754; Demerara; Jenman 4547 as to plant with appressed short pubescence. FRENCH GUIANA: Route de Cayenne Rochambeau, Halle 434. VENEZUELA. Bolivar: Paso de Cardozo, 10 km S of Ciudad Bolivar, alt 50 m, Killip 37695; Anzoategui: bosque de El Riecito al sureste de Santome, alt 240 m, Pittier 14557; Sucre: Peninsula de Paria, Cerro de Rio Arriba, alt 600-700 m, Steyermark & Rake 96271. BRAZIL. Amapa: along road to Amapa, vicinity of km 108, Rio Pedreira, Pires & Cavalcante 52216; road to Amapa, km 134, Agua Azul, Pires & Cavalcante 52275.
The Brazilian specimens appear somewhat intermediate between C. tontanea and C. guianense. Many of the specimens cited above have been previously identified as C. guianense.