Monographs Details: Symplocos pycnophylla Sleumer
Authority: Maguire, Bassett. 1978. The botany of the Guayana Highland--part X. Mem. New York Bot. Gard. 29: 1-288.
Family:Symplocaceae
Description:Distribution and Ecology - Distribution. Shrubs or small trees to 7 m, confined to the uplands of Guayana. VENEZUELA. Estado Bolivar: Rfo Cuquenan, Feb 1910, Ule 8730, (holotype B , not seen by us); shrub 10-15 ft tall, Mt. Roraima, 2255-2620 m alt, 27 Sep 1944, Steyermark 58761 (isotype of S. roraimensis, NY); small tree 2 m high, flowers yellowish, rocky elevations, occasional, Kavanayen Savannas, 1300 m alt, 30 Mar 1952, Maguire 33681 [male] (NY, VEN); bushy tree 5-7 m, frequent at 2050 m alt, talus cloud forest, Churi-tepui, Chimanta Massif, 30 Jan 1953, Wurdack 34271 (NY, US, VEN, F, K , U). Territorio Amazonas: bushy tree 3 m high, flowers white, apparently [male], stamens variable in number, west rim, Cerro Paru, 2000 m alt, 7 Feb 1951, Cowan & Wurdack 31302 (NY, US, F, VEN, U); tree 6 m, on escarpment at 2000 m alt, Cerro de la Neblina, 12 Jan 1954, Maguire et al 37246 [female] (NY, VEN, US, K, P, MO, GH).

Discussion:1. Symplocos pycnophylla Sleumer, ex descr. Fedde Rep. Sp. Nov. 42: 266. 1937. Fig 108. Symplocos roraimensis Steyermark, Fieldiana Bot. 28: 494. 1953. Type. Shrub 10-15 ft tall, calyx olive green, petals pale green, SW face Mt. Roraima, 2255-2620 m alt, Estado Bolivar, Venezuela, Julian A. Steyermark 58761 (holotype F, isotypes NY, VEN). Type. Am oberen Rio Cuquenan (in der Nahe des Mt. Roraima), Strauch mit weissen Bluten, II. 1910 (E. Ule no. 8730, typus in Herb. Berlin). We have had opportunity to examine specimens of Symplocos roraimensis Steyermark, the near-related S. bogotensis Brand (isotype COL, 3000 m, Triana sn, in 1851-1857, NY), and a number of more recently collected specimens of my own and those of colleagues. Reluctantly we are required to consider the Guayana materials here cited as the same species, all of which from the description and type locahty must be referred to the older name S. pycnophylla. These collections are held together by consistent floral characters and vegetative facies. Our species may be distinguished from Symplocos bogotensis, in which the corolla lobes twice exceed the calyx lobes, whereas in S. pycnophylla the calyx and corolla are essentially equal in length. An examination of pollen of Symplocos pycnophylla {Cowan & Wurdack 31302), Fig 109A-C, shows the grains of this species to be oblate in equatorial view and semi-triangular in polar view, triaperturate with prominent pores, consisting of relatively short, slit-like openings at distal ends of each of the triangle apexes. The tectum is strongly globulate (as defined by van der Meijden, Pollen Morphology of Indo-Pacific Symplocos, Pollen et Spores 12: 513-551. 1970) and seems to fall within his 2-3 Glauca subtype. Van der Meijden states that eight American species of the Brand subgenus Epigenia examined by him have pollens of this subtype. Symplocos pycnophylla as interpreted here does indeed belong to the subgenus Epigenia as defined by Brand.^13 So far as observed in the available material, our plant is wholly dioecious. Symplocos roraimensis must be placed in synonymy under S. pycnophylla (see under this species).
Distribution:Venezuela South America|