Achlyphila

• Authority

  Carlquist, Sherwin J. 1960. Anatomy of Guayana Xyridaceae: Abutboda, Orectanthe and Achtyplola. Mem. New York Bot. Gard. 10: 63-117.

• Family

  Xyridaceae

• Scientific Name

  Achlyphila

• Discussion

  In respect of exodermis and cortex structure, the root of Achlyphila distich a (fig. 7) conforms to the pattern outlined above. The endodermis, which stains bright orange-red (rather than pink, as in Abolboda) with safranin, possesses U-shaped thickenings not exactly like any mentioned above. The endodermis may be two cells wide in places, although this condition is rare and localized. Two roots examined proved to be 11-arch and 13-arch respectively. No central vessels are present. This is surprising in view of the large diameter of the vascular core. Instead, one to several cells that have the wall-thickening and staining characteristics of endodermis cells (figure 7, dark area in center of vascular core) are present in this position. On account of their characteristics, they probably should be regarded as endodermal cells, despite their location. Phloem groups are extremely small and are scattered among the sclerified cells of the vascular core. The pericyele is not interrupted. Significantly, there are two or three layers of cells (as compared to fewer in Abolboda) between the pericyele and the outermost vessels. Occlusion by tannin-like materials is characteristic of many vessels.