Monographs Details: Brachymenium notarisii (Mitt.) A.J.Shaw
Authority: Buck, William R. 1987. Bryostephane Steereana: A Collection of Bryological Papers Presented to William Campbell Steere On The Occasion of His 80th Birthday. Mem. New York Bot. Gard. 45: 1-749.

Bryum notarisii Mitt., J. Linn. Soc. Bot. 8: 3. 1865; B. semicompletum Mitt., J. Linn. Soc. Bot. 8: 3. 1865, nom. nud. in syn.; Mielichhoferia notarisii (Mitt) Jur., Bot. Zeit. 24: 178. 1866; Haplodontium notarisii (Mitt.) Broth., Nat. Pfl. 1(3): 590. 1903. Syntypes: MADEIRA. "On the earth in moist places, Johnson s.n.; in insulis freti Bonifacii maris Mediteranei, De Notaris s.n." (lectotype NY!).

There are two large envelopes on one sheet (among several other sheets) representing this species in the Mitten herbarium. The top envelope contains the type collected by J. Y. Johnson, present in triplicate. It is labelled B. semicompletum and also includes a latin diagnosis and some pencil drawings. The bottom envelope contains two more isosyntypes, one of which is the lectotype. According to data on the herbarium labels, J. Y. Johnson, a zoologist w h o spent winters in Madeira, collected B. notarisii on "banks near Funchal."

Mitten (1865) considered B. notarisii as closely related to B. alpinum With., and indeed, there is some gametophytic resemblence (lanceolate or ovate-lanceolate leaves, long, narrow leaf cells, etc.). The species is, however, recombined here in Brachymenium. The placement is made largely in consideration of traditional generic concepts in the Bryaceae. Brachymenium diflfers from Bryum in having the endostome highly reduced and generally lacking diflferentiated segments. On the other hand, several species of Bryum, for example, B. wrightii Sull., have very reduced endostomes and Brachymenium is, in my opinion, poorly defined by peristome characters alone. Extreme examples of peristome reduction occur in almost every genus of the Bryaceae. Pending the development of a refined concept of Brachymenium, B. notarisii is placed there; it certainly does not belong in the Mielichhoferioideae. The complete lack of an endostome is unusual in Brachymenium, but the diflference is only one of degree when compared to Brachymenium species that have a basal membrane only.