Fissidens kurandae Broth. & Watts, Proc. Linn. Soc. N.S. Wales 43: 550. 1918, syn. nov.
Fissidens holstii has a hyaline dot in the laminal
cells and marginal cells occasionally appear
thickened because of a tilting at the edge (sometimes
seen in the two new species) or are vaguely
and sporadically bistratose due to an oblique division
in marginal cells but not conspicuously
bistratose as in F. nitens where the thickened
border is 2-4 cells wide. Fissidens nitens and F.
holstii, judging by their areolation and costal
structure, are clearly unrelated to F. sufflatus and
Fissidens javanicus Dozy & Molk., an Asian
species, also has thickened leaf margins, resembling
F. sufflatus in the swollen unistratose margin
of the vaginant lamina but diflfering from
both F. sufflatus and F. pseudopallidus in several
features, in particular the bistratose or tristratose
apical and dorsal laminal margins and the extremely
large hyaline axillary nodules on the stem,
characters previously reported by Fleischer
(1904), and Iwatsuki and Suzuki (1982).
Fissidens pallidus Hook. f. Sc Wils., an Australian
and New Zealand species, has a wide distribution
in eastem Australia extending from
Tasmania to north Queensland where it is not uncommon on earth banks in rainforest and
sometimes occurs in association with F. pseudopallidus.
Fissidens pallidus (Figs. 62-67, drawn
from type: NEW ZEALAND. North Island,
Colenso 391, bm) appears to be very closely related
to F. pseudopallidus having sporophytic
similarities and resemblances in general habit,
areolation and costal structure although the vaginant
lamina (Fig. 62) is generally proportionately
longer, usually about two-thirds leaf length
instead of half or less and the leaf apex usually
less tapered (Fig. 63). The most striking differences,
however, are in the characters ofthe marginal
cells; in F. pallidus the marginal cells (Figs.
64-67) ofthe apical and dorsal laminae are not
swollen as in F. pseudopallidus and the vaginant
lamina, unlike that of F. pseudopallidus in surface
view lacks a broad pale border which, in
cross section, is not thickened but tapers to the
margin. The spores of F. pallidus are smaller,
7.5-11 µm, than those of F. pseudopallidus which
are 12-15 µm.