Monographs Details: Hamelia patens Jacq. var. patens
Authority: Elias, Thomas S. 1976. A monograph of the genus Hamelia (Rubiaceae). Mem. New York Bot. Gard. 26: 81-144.
Family:Rubiaceae
Description:Distribution and Ecology - Specimens examined. UNITED STATES OF AMERICA. Florida: Brevard: Indian River region, hammock, Fredholm 5587 (GH, M O , US), Curtiss 1128 (A, F, GH, MO, NY); near Georgiana Landing, Canby s.n. (NY). Colher: Pinecrest, S of Tamiami Trail between mile 40 & Ochopee, Field & Lazar s.n. (F, US); Big Cypress Swamp, rocky soil along roadside, Correll 5965 (GH); Miles City, Brass 15758 (GH, US); W of Ochopee, Cooley et al. 6177 (NY). Dade: Biscayne Bay, old field near Cutler, Curtiss 5500 (F, GH, MO, NY, NYS, US); Palmer 221 (GH, MO, US); S of Cutler, Black River region, Eaton 271 (F); Cutler's Hammock, Sargent s.n. (A); in hammocks between Coconut Grove & Cutler, Small & Carter s.n. (F), 1193 (F, NY); Addison Hammock, ca. 5 mi S of Coconut Grove, Harris C16 119 (F, GH); hammock between Miami & Coconut Grove, Small & Small 4661 (NY); hammock near Coconut Grove, Young 219 (US); hammocks near Miami, Small & Carter 809 (NY), 4540 (NY), s.n. (GH); Buena Vista, dry sandy soil, Moldenke 316 ( MO, NY); hammock, 25 miles W of Miami, Moldenke 3770 (NY); Brickell Hammock, O'Neil s.n. (MO); Goodbum Hammock, Small & Mosier 5921 (GH, N Y , US); Hattie Bauer Hammock, Small et al. 6425 (MO, NY), Alexander s.n. (NY); Sykes Hammock, Small et al. 5792 (NY); without exact locality, Eaton 826 (F). Highland: Sebring, hammocks, Vinten 61 (NY), McFarlin 8266 (F); Highlands Hammock State Park 6 miles W of Sebring, Wilbur & Webster 2600 (GH, NY). Lake: vicinity of Eustis, hammock, Nash 1278 (A, F, G H , M O , NY, US). Manatee: vicinity of Manatee, Simpson 48 (F, US). Marion: Dallas, Cooper s.n. (GH). Monroe: Cape Sable, sand dunes. Small 7698 (NY): N Key Largo, hammock. Stern 1449 (US), Cooley et al. 9269 (GH); Key Largo, hammock. Brass 20491 (US), Sargent s.n. (A), Janish & Janish 491 (MO); Sugar Loaf Key, Pollard et al. 27 (F, N Y, US): Big Pine Key, Killip 44398 (GH), Killip & Swallen 40339 (F), Stern 1496 (US), Small 8152 (NY), Robertson 4 (GH), Brizicky et al. 421 (A); Key West, Blodgett s.n. (NY), RugeUll (F, GH, MO, NY, US). Orange: at Clarcona, Pieters 96 (F, US), Meislahn 113 (US); Lake Apopka, O'Neill s.n. (MO). Palm Beach: Craigin Estate (cultivated), Lidestrom 4272 (MO). Seminole: cuUivated, Schallert 16059 (WIS). Sumter: without exact locality. Smith s.n. (US). Without locality, Garbers.n. (F, GH, NY, US), Chapman s.n. (NY). BAHAMAS. New Providence: Grantstown, P. Wilson 8169 (F, NY), Wight 147 (F, GH, NY); Nassau, Northrop & Northrop 40 (F), Hitchcock s.n. (MO); Lake Cunningham, Britton & Brace 635 (F). CUBA. Camaquey: vicinity of La Goria, Shafer 116 (NY). La Habana: near Havana, Palmer & Riley 1142 (US), Day 293 (NY); Batabano, Shafer 119 (F); San Antonio de Los Baiios, Curtiss s.n. (GH), Shafer 35 (F); Santiago de las Vegas, Hermann 611 (F, MO, NY); Rib Almendares to Playa de Marianao, P. Wilson 9462 (NY). Las Villas (Santa Clara): Cienfuegos, Cienequita, Combs 29 (F, G H , M O , N Y ) ; Soledad, Hodge et al. 4038 (GH, MO, NY, NYS, US), Hunnewell 11553 (GH), Jack 4155 (US), 4671 (A), 4789 (A); Rib San Joquin, Howard 5118 (GH, N Y ) ; rocky hill 0.2 miles W of Rib San Juan Crossing to Trinidad, Howard et al. 383 (A); Gavilan, roadside fields & copses. Smith & Hodgdon 3186 (F, GH, MO); San Bias, Rehder 1151 (A); above San Bias, Sierra San Juan, Morton 4047 (US); El Junco region above Siguanea, Howard etal. 204 (A); Sierra Trinidad, Las Vegas de Matagua, Morton 10818 (US); Hanabanilla Falls, Britton et al. 4832 (NY); Siguanea, 400 m Britton & Wilson 4892 (NY); Caibarien, pastures, Ansovin 657 (GH). Matanzas: E of Matanzas, Britton et al. 185 (NY); Playa Matanzas, Britton et al. 35 (NY). Oriente: Baracoa to Rib Guaba, Shafer 4319 (NY); vahey of Rib Bayamita, S slope of Sierra Maestra, Maxon 3938 (US); El Cobre, Santa Elena Estate, Clemente 7582 (GH, N Y , US); near Monte Verde, Wright 227 (GH); valley of Rib Matamoros, S of Holguin Shafer 1372 (NY); N spur of Sierra Maestra W of Rib Yao, Morton & Acuna 3445 (US); Punton de Cuero, S coast of Oriente, Morton & Alain 8855 (US); la Playa de Siboney, Lopez 169 (US); near Santiago, Clemente 1818 (GH), Havard $ (NY), Taylor 404 (NY). Pinar del Rib: near Guanajay, Palmer & Riley 663 (US); Bafios San Vicente, Britton et al. 7488 (NY); vicinity of Mendoza, Shafer 10590 (MO, N Y , US); Sierra de Anafe, P. Wilson 11494 (NY). Province unknown: without exact locahty, Jacquin s.n. (F frag ex Vienna). Isle of Pines: Upper Rib de Los Indios, Jennings 673 (NY); Pedernales Point, Millspaugh 1412 (F); without exact locality, Taylor 212 (F, G H , N Y , US). JAMAICA. Quarry near Spanish Town, Harris 12403 (F, G H , M O , N Y , US); bank of Rib Cobre, near Spanish Town, Britton 3108 (NY); Constant Springs, gully, Philipson 549 (A); near Widcombe, roadside thicket, ca. 330 m. Proctor 10633 (GH). HAITI. vicinity of Anse Galette, Gonave Island, Leonard 3202 (US); from Anse d'Azure to 3 mile W of Jeremie, Siebert 1722 (MO); Bayeux, near Port Margot, rocky hillside, Nash 6 (F, NY); vicinity of Miragoane, Everdam 595 (GH, US); between Mornes a Cabrites & Mirebalais, Holdridge 1159 (NY, US); Massif de la Salle, Port-au-Prince, Ekman 6614 (A, N Y , US), Rose 22100 (GH, N Y , US), Holdridge 1018 (US); Massif de la Hotte, between Trou-d'Enfer & La Seringue, E k m a n 10850 (A, N Y , US); vicinity of Port de Paix, Leonard & Leonard 11156 (A, GH, US); vicinity of La Vallee, Tortue Island, Leonard & Leonard 11230 (US, NY). DOMINICAN REPUBLIC. Barahona, Fuertes 48 (F, G H , M O , US); S W of Barahona, Canada Maluca, Howard 12202 (A, US); Boca Chica, Augusto 1400 (NY); Consuelo, edge of forest, Taylor 121 (NY); Guayacanes, Augusto 1540 (NY); Haina, Paris 295 (US); La Ceiba Demao, Valeur 493 (F, M O , US); Montiada Nueva, forested hillslopes S E of Polo, 1200 m, Howard & Howard 8422 (F, G H , N Y ) , 8451 (F, G H , N Y , US); near Ocoa, Turckheim 3643 (NY, US); trail between Pedernales & Aceitial, 1200 m , Howard & Howard 8245 (F, G H , N Y , US); beach forest at Sosua Settlement, Holdridge 513 (NY), 514 (NY); 20 km W of San Pedro de Macoris, Howard & Howard 9491 (F, G H , N Y , US); Santo Domingo, Millspaugh 823 (F), Rose et al. 3757 (GH, N Y , US); near Santo Domingo at Rib Ozuma, Turckheim 2842 (NY), E k m a n 12323 (US); vicinity of Santo Domingo, 0-25 m, Allard 13318 (US), 13438 (US), 15913 (US); road to Valverde, Valeur 703 (A, F, MO, US). Locality unknown: Canby s.n. (NY). PUERTO RICO. Arecibo, hillsides. Heller & Heller 339 (F, N Y ) ; Aibonito, Otero 290 (MO); Cayey, Urban 2476 (F, M O , US); vicinity of Mayaguez, Britton & Marble 674 (NY), Otero 545 (MO), Urban 65 (NY, US); wooded area on route 191 on S side of Sierra Luquillo, Howard 16604 (GH, M O ) ; near Juan Diaz, Pena de las Cuevas, Britton & Marble 2315 (NY); 4 miles from Ponce, on Adjuntas road. Heller 6083 (A, M O ) ; arroyo W of Ponce, Britton & Britton 7252 (GH, N Y ) ; San German, Miller 1665 (US); near San Sebastian, roadside, 200 m , Liogier 9928 (GH, N Y ) ; without exact locality, Bertero s.n. (MO), Urban 2621 (NY). VIRGIN ISLANDS. St. Croix: Crequis, Ricksecker 351 (F, G H , M O , NY, US), 442a (F, M O , US), Thompson 226 (GH); Mt. Steward, Howard 15287 (A). St. Thomas: without exact locality, Eggers s.n. (MO), Eggers s.n. (US). GUADELOUPE. near Pointe Noire, Duss 4124 (F, N Y , US); Abymes, Questel 698 (US); near Mome-a- I'Eau, Stehle 1819 (A, US). DOMINICA. without exact locality. Hooker s.n. (NY). MARTINIQUE. at site of old botanic garden at St. Pierre, Stehle 2145 (NY), 4556 (US), E w a n 17078 (US); Lamentin, Fairchild3764 (A, US); without exact locahty, Duss 953 (NY). MEXICO. Campeche: Tuxpeiia, Lundell 905 (F, GH, MO, NY, US). Chiapas: Escuintla, Matuda 653 (F, US); M T . Ovando, 900 m, Matuda 4203 (A, F, GH, NY, US); Esperanza, Matuda 16773 (F); near Tuxtla Gutierrez. Carlson 2151 (F); ca. 9 miles E of Tuxtla Gutierrez, King 2764 (NY); 22 k m N of Tuxda Gutierrez, 1500 m , Breedlove 10673 (F); near Tenejapa, 1550 m, Breedlove 6132 (F); San Bartolome, Kaplan 211 (F); E of Ocozocoautla, dry rocky hills, Moore 2522 (GH), Seler 2118 (GH); along road between Teneapa & Yajalon, 1000-1650 m. Nelson 3259 (US); Palenque ruins, Seler 5477 (GH, US); Jalisco, Purpus 9210 (F, GH, MO, US). Colima: Armeria, Palmer s.n. (GH); Hacienda Colomo, 18 k m S of Manzanillo, open mixed forest. West 3530 (GH). Guerrero: Acapuico, on Pan American Highway, Frye & Frye 2566 (GH, MO, NY, US); La Correa, 50 m, Langlasse 439 (GH, US); Montes de Oca, Petatlan, Hinton 10327 (NY, US). Hidalgo: Chapulhuacan, Kenoyer 821 (MO); below Chapulhuacan, 700 m, Moore 2880 (GH); vicinity of Huejutla on road to San Felipe Orizatlan, Moore 2202 (GH); Santa Maria, vicinity of Santa Ana, Quintero 1631 (WIS); Santa Ana, Frye & Frye 2653 (GH, M O , N Y ) . Mexico: vicinity of Rancho del Flores near San Pablo, 2430- 2520 m, Happ 288 (MO). Nuevo Leon: E of Santiago, Gregg 1062 (MO), 1071 (MO). Oaxaca: Cuicatlan, Smith 7 (GH); Chiltepec & vicinity, ca. 20 m, Calderon 273 (A, US); near Totontepec, 1250-1800 m. Nelson 787 (US); ca. 2 k m N of Ixhuatan, open grazed areas. King 1994 (NY, US); plains N of Tehuantepec, Alexander 96 (NY), 97 (NY). Puebia: vicinity of Huauchinango, 600 m , Froderstrom & Hulten 1057 (A, F, NY). Queretaro: 74 miles N E of Zimapan, Waterfall & Wallis 14255 (US). Quintana Roo: Cozumel Island, Millspaugh 41 (F, GH); vicinity of San Miguel Cozumel, Lewis 6855 (MO). San Luis Potosi: Las Canoas, limestone cliff, 900-1000 m , Pennell 17949 (F), near Tamazunchale, Edwards 466 (F, MO, NY), Barkley 17M093 (F); 15 miles N of Tamazunchale, Hitchcock & Stanford 6914 (NY); 48 miles N of Tamazunchale, Waterfall & Wallis 14293 (US); near Axtla, Weaver 656 (GH, US); El Banito, 33 m. Chase 7557 (F, GH, MO); valley of Rib Tampaon, 33 m. Chase 7515 (F, GH, MO, NY); vicinity of El Salto above El Naranjo, Duke 3921 (MO); El Salto, Quintero 117 (WIS); S of Valles, Leavenworth 212 (F, GH, MO, NY), Frye & Frye 2961A (GH); Valles, 233 m. Vines 3323 (US); Tomasopo canyon, Pringle 3677 (F). Sinaloa: near Coacoyolitos, Ortega 5869 (MO). Tabasco: 11 k m up Rib Mezcalapa above San Manuel, Gilly & Hernandez 110 (GH); near Rib Teapa, 1 k m N of Teapa, Gilly & Hernandez 218 (GH); HuimanguiUo, savanna region, Barlow 30/158 (WIS); Balancan, Matuda 3245 (A, F, NY). Tamaulipas: Galeana Cafion, Kenoyer & Crum 3499 (A); dry shady banks of Rib San Marcos, 6 miles W of Cuidad Victoria, 460 m. Meyer & Rogers 2480 (GH, MO); near Ciudad Victoria, Happ 24 (MO), Kenoyer & Crum 3288 (A), Palmer 63 (F, GH, MO, NY, US), 517 (F, GH, MO, NY, US); 10 k m N W of El Progreso, Stanford et al. 1046 (GH, MO); vicinity of Tampico, ca. 15 m. Palmer 184 (GH, MO, NY, US), 576 (GH, MO, NY, US); at La Noria, Wooton s.n. (US); El Mante, Kenoyer C124 (GH). Tres Marias Islands: Maria Madre, Howell 10470 (F, US). Vera Cruz: ca. 20 miles S of Acayucan, King 2734 (NY); Atoyac, Matuda 1398 (A, F, MO); Boca del Rio, King 2691 (NY, US); Chinameca, Orcutt 3164 (F, GH, MO, US); Cordoba, Fisher 35451 (F, MO, NY, US), Bourgeau 1607 (US), Matuda 166 (F), Plunkett 209 (F); Fortin, White 311 (US); Cerro Huacapan to 2 k m N of Atzacan, 1350 m , Rosas 361 (GH, MO); Jalapa, Barkley et al. 2621 (F), Orcutt 3164 (F, GH, MO, US), Coulter 202 (GH); Lobos Island, low sand cay, Mclntire 6L (WIS), Chavez s.n. (WIS); Montepio, 19 k m E of Catemaco, Quintero 2206 (WIS); near Motzorongo, Nelson 161 (US), Orizaba, Fisher 166 (US), Bourgeau 2627 (GH, US), Balls 4324 (A), Smith 459 (MO), Bilimek 191 (US), Miller s.n. (NY); Cerro Escamela N of Orizaba, Rosas 331 (GH, MO); vicinity of Pueblo Viejo, Palmer 425 (GH, MO, NY, US), 538 (F, GH, MO, NY, US), 546 (GH, MO); Panuco, Seler 680 (GH, US); Papantla, Seler 3671 (GH, US); 30 k m SE of Poza Rica, Romero 36 (WIS); near barranca of Texolo, 1150 m , Pringle 8150 (F, GH, MO, NY, US); Cerro San Cristobal, 0.5 miles S of Orizaba, Manning & Manning 53798 (GH); 5-10 km N of Tuxpan, Boke & Floratos 19 (GH); without exact locahty, Muller s.n. (GH, NY). Yucatan: Chichen Itza, Bequaert 6 (A, F), Millspaugh 122 (F, GH), Steere 1002 (F, M O ) , Dzitas, Stewart 308 (GH); Izamal, Gaumer s.n. (F); Mani, Seler 3915 (A, G H , F); Merida, Schott 174 (F, US); Kabah, near Mayan ruins, Lewis 6898 (MO); La Vega, Goldman 637 (US); San Anselmo, Gaumer 1929 (F, GH, MO); Tizimin, Swallen 2527 (US); Kancabdzonot, Gaumer 23621 (F, G H , US); without exact locality, Gaumer 727 (A, F, GH, MO, NY, US), 24067 (F, GH, MO, US). GUATEMALA. Alta Vera Paz: Coban, Hunnewell 17258 (GH); Cubilquitz (?), Turckheim 222 (F, GH, MO, NY); Sesisp, Turckheim 750 (US); near Finca Sapacuite, Cook & Griggs 762 (US); Finca Trece Aguas, Goll 22 (US); Pantin, below Tamahu, ca. 600 m,Standley 70864 (F). Chiquimula: between La Laguna and Chiquimula, 500-1000 m, Steyermark 30734 (F). Escuintla: El Baul, Tonduz 101 ( MO, US); near Escuintla, ca. 135-300 m, Standley 63949 (F), Smith 1974 (GH, NY, US); between Rib Jute and Rib Pantaleon, 540-720 m, Standley 63422 (F); Santa Lucia, Degener & Degener 26481 (US); Anubis, near Obispo, Muenscher 12376 (F, GH); without exact locality, Aguilar 1664 (F). Huehuetenango: between Finca San Rafael and Finca Providencia, 800 m, Steyermark 49550 (F); without exact locality, Seler 2654 (GH, US). Jalapa: Laguna de Ayarza, Heyde & Lux 3976 (US). Jutiapa: Agua Blanca, Tyada 49 (US). Izabal: vicinity of Quirigua, in marsh or moist thicket, Standley

Distribution and Ecology - 23809 (GH, US), 23850 (GH, US); Rib Dulce, Livingston, Smith 1788 (US); between Los Amates and Quirigua, 80-83 m, Steyermark 38314 (F). Suchitepequez: S of Alotenango Farm, 30-50 m, Steyermark 47792 (F); vicinity of Mazatenango, Bernoulli 536 (NY), Maxon & Hay 3453 (US); vicinity of Tiquisate, 100 m, Steyermark 47676 (F). Peten: Carmehta, Egler 42-209 (F); La Libertad and vicinity, Aguilar 10 (A, MO), 39 (A), Lundell 3103 (F); Plaza Mayor Tikal ruins, Molina 15814 (F); Uaxactun, Cook & Martin 148 (US); 35 k m from St. Elena, Molina 15463 (F). Quezaltenango: Colomba, 850 m, Skutch 1283 (A, F, NY, US). Quiche: without exact locality, Aquilar 1138 (F). Retalhuleu: N of Nueva linda, ca. 120 m, Standley 87748 (F); San Felipe, 500 m, Steyermark 34506 (F); Rib Samala, Shannon 592 (A, F); Ocos, 1-2 m, Steyermark 37781 (F). Santa Rosa: S of S of Los Cerritos, on road to El Ahumado, Standley 79697 (F); Santa Rosa, Heyde & Lux3159 (GH, NY, US). Solola: Rib Bravo, Shannon 391 (US); Santa Barbara, Shannon 130 (US). Zacapa: Gualan, 140 m, Deam 269 (F, GH, MO, NY, US), 6290 (A, F, GH, N Y ) , Kellerman 5632 (US); trail between Rib Hondo and waterfalls, 250-400 m , Steyermark 29408 (F). Province unknown: without exact locality, Heyde 325 (US). BRITISH HONDURAS. Belize: Turneffe Island, Egler42-39{F); Lighthouse Reef, Half M o o n Cay, Stoddart 138 (US). El Cayo: Begonia Hill, 8 miles E of Central Farm, Dwyer et al. 149 (MO, SIU); road to Mai Fire Lookout Station, 400-530 m, Dwyer et al. 184 (MO, SIU); S W of Cayo, summit of Mayan Ruins, Dwyer et al. 250 (MO, SIU); river bluffs, Bartlett 11439 (A, F, M O ) ; near Agustin, Molina 216(F). Corozal: without exact locality. Gentle 227 (F, US). Orange Walk: Honey Camp, Lundell 15 (US), 53 (F), 132 (US), 498 (A, F, GH, NY, US); vicinity of Tower Hill Estate, Egler 42-97 (F), Karling 24 (A, F, GH, US), Karling 37 (US); without exact locahty, Winzerling I-ll (US). Stann Creek: Big Creek, 33 m, Schipp 105 (F, GH, MO, NY, US); along Hummingbird Highway, N of Stann Creek Town, Dwyer et al. 425 (MO, SIU); Middlesex, Record s.n. (US); Tipperary Road to Silk Grass, Stevenson 3 (US). EL SALVADOR. Ahuachapan: vicinity of Ahuachapan, Standley 19942 (GH, N Y , US); without exact locality, Padilla 150 (US), 151 (US), 364 (US). La Libertad: vicinity of Ateos, Standley 23383 (US). La Union: vicinity of La Union, Standley 20834 (GH, US). San Miguel: Hacienda Potrero Santo, S side of Lake Olomega, Tucker 830 (F, US); vicinity of San Salvador, Calderon 157 (GH), 424 (US), Standley 19101 (GH, US), Renson 125 (NY); along road from San Martin to Laguna de Ilopango, Standley 22499 (GH, US). San Vicente: vicinity of San Vicente, 350-500 m, Standley & Padilla 3795 (F), Standley 21209 (GH, US). Sonsonate: Rib Acachapa near Hacienda Las Tablas, ca. 780 m. Tucker 1361 (F, US); vicinity of Izalco, Standley 21838 (GH, US); roadside between Sonsonate and Izalco, Porter 1272 (GH); vicinity of Santa Emila, Standley 22129 (GH, US); vicinity of Sonsonate, Standley 22315 (US). HONDURAS. Atlantida: San Alejo, Mallery 5019 (NY); near Tela, Lancetilla Valley, Standley 53272 (A, F, US), 54451 (A, F, US); in thicket between Tela and Lancetilla, Yuncker 4497 (A, F, MO); in thicket on plains near Ceiba, Yuncker et al. 8431 (F, GH, MO, NY, US); trail W of Rib Tela, P. Wilson 109 {NY, US). Central District: Tegucigalpa, Hagen & Hagen 1260 (NY). Comayagua: Las Lima, 1000 m, Edwards 238 (A, F); near Minas de Oro, Molina 8469 (F); Pito Solo, Lake Yojoa, Edwards P-434 (A, F); vicinity of Siguatepeque, ca. 1050 m , Standley & Chacon 6905 (F), Yuncker et al. 5672 (F, GH, MO), 6312 (F, GH, MO); between Trincheras and Montanuelas, Molina 10858 (F). Copan, 5 k m S of Santa Rita, Molina 6659 (F), vicinity of ruins of Copan, 660 m , Carlson 527 (F); Las Amates to Hacienda El Limon, Blake 7334 (US). Cortes: near Choloma, San Pedro Sula, Molina 6670 (F), Cuyamel, Record & Kuylen 9951 (NY, US); Rio Lihdo near Lake Yoja, 300 m , Williams & Molina 11397 (A, F, MO); 5 k m from San Pedro Sula, Molina 3815 (F); Rib Ulua, Pimienta, Molina 5644 (F). Lempira: near Gracias, Molina 6571 (F). Morazan: Agua Amarilla, pine-oak forest, Williams & Molina 10704 (F, GH, MO); Aldea de Suyapa, Quebrada Suyapa, Molina 2571 (F, G H ) ; El Jicarito, above El Zamorano, 820-950 m, Standley 20781 (F); along Rib de la Orilla near Cerro Majicaran, 800 m, Williams 14061 (F, G H ) ; slopes of Mt. Uyaca, 1400 m, Williams & Molina 10196 (A, F, M O ) ; drainage of Rib Yeguare, Glassman 1822 (F, NY, WIS); vicinity of El Zamorano, 780-900 m, Standley 3849 (F); Zamorano, 950 m, Williams & Molina 11939 (F, GH, MO). El Paraiso: along Rib California, Sierra de la Villa Santa, 700 m, Williams & Molina 10478 (A, F, MO); near Yuscaran ca. 960 m, Standley et al. 1204 (F); drainage of Rio Yeguare, Molina 3958 (F, G H ) . Olancho: near Escuela Granja Demonstrativa de Catacamas, 400 m , Molina 8320 (F). Santa Barbara: near San Pedro Sula, Thieme 5271 (F). Valle: Tigre Island, vicinity of Ampala, West 3541 (GH), Standley 20718 (US); Disposicion Island (?), Stork & Worth 8863 (A). Yoro: near Progreso, Hottle 75(F). NICARAGUA. Chinandega: Ameya, Maxon et al 7134 (US); Chinandega, Baker 2002 (GH, M O , US). Granada: Volcan Mombacho, Baker 2478 (GH, M O , US). Jinotega: Ocotillo near St. Lastenia, 1550 m , Williams et al. 27846 (F). Managua: Casa Colorada and vicinity, ca. 300-800 m , Maxon et al. 7367 (NY, US), 7413 (US), Standley 8437 (F), Sierra de Managua, 400-900 m. Gamier A165 (US), A633 (US), 978 (US), Grant 1006 (A, F); Managua, Chaves 196 (US); Las Nubes and vicinity, S of Managua, Maxon et al. 7533 (US); Ometepe Island, Lake Nicaragua, Shimek & Smith 46 (F); near San Jacinto, Williams & Molina 10965 (F, G H , M O ) . Masaya: S W slopes of Santiago Volcano, near Masaya, M a x o n 7687 (US). Matagalpa: ca. 15 k m N E of Matagalpa along Rib Las Canas, Williams 27513 (F); between Matagalpa and Jinotega, cloud forest, Williams et al. 23310 (F), 24650 (F). Zelaya: region of Braggman's Bluff, Englesing 161 (F). Province unknown: without exact locality, Wright s.n. (MO, NY, US). COSTA RICA. Alajuela: hills 2 k m S of San Ramon, Fosberg & Brenes 27360 (F). Cartago: vicinity of Cartago, Cooper 5789 (US), 5790 (GH, US), Standley 33400 (US), Stork312 (US); Cerro de la Carpintera, Standley 34523 (US); Chitana, Rojas 204 ( M O ) ; vicinity of Santiago, Maxon 78 (US); along stream near Tres Ribs, Williams 16135 (F); near Santa Maria, Tonduz 11660 (US); near San Ramon, Brenes 14496 (GH); vicinity of El Alto Railroad Station, Allen 660 (MO); vicinity of Santiago, 1050 m, Maxon 78 (NY, US). San Jose; Asseri, Hunnewell 16759 (GH); Chitaria, Solis 204 (F); vicinity of El General, Skutch 2410 (A, GH, MO, NY, US), 4189 (US), 4319 ( M O ); vicinity of Escasu, ca. 1250 m, Standley 32317 (US); near Guadalupe, Tonduz 1545 (GH, US), 1784 (US), 7127 (US); vicinity of Santa Maria de Dota, Standley 42096 (US), Standley & Valerio 43134 (A, US); San Pedro, Greenman & Greenman 5316 ( M O ); between San Pedro de Montes de Oca and Curridabat, ca. 1200 m, Standley 32832 (US); vicinity of San Jose, Holway 390 (US), Biolley 65 (F), Tonduz 8016 (US); vicinity of San Sebastian, ca. 1160 m , Standley 49310 (A, US); Rib Tiribi, Alfaro 33986 (US). Puntarenas: vicinity of La Presa, ca. 3 miles above Palmer, Allen 5283 (US); Boruca, Tonduz 4665 (US). Province unknown: without exact locality, Kuntze 2148 (NY); Calamanca (?), Tonduz 8633 (US), 8722 (US). PANAMA. Canal Zone: near Albrook Air Force Base, Gallegos et al. 5146 (MO); Balboa, Standley 25244 (F, US), 26072 (US), 27131 (US), Rowlee & Rowlee 448 (NY, US); Chiva Chiva Trail, near Miraflores Lake, Tyson 1400 (MO), 1404 (MO); vicinity of Cocoli, Tyson 1635 (MO), Stern et al. 308 (GH, M O , US), 347 (GH, M O , US); secondary growth on hillside below Contractor's Hills, Stimson 5242 (MO); near Survival School at Curundu, Tyson 1073 (MO); Farfan beach, Kirkbride 80 ( M O ) ; Paraiso railroad station, Hayes 38 (NY); near Madden Dam, Dwyer & Hayden 4 (MO), Dwyer & Robyns 11 (MO), Dwyer & Elias 7487 (GH, MO, US), Ebinger 847 (MO), Woods et al. 5085 (DUKE, MO), Boy Scout Camp road, Kirkbride 40 (MO), 41 (MO); vicinity of Miraflores Lake, White & White 188 (GH, MO, US); without exact locahty. White 122 (F, G H , MO). Chiriqui: vicinity of Boquete, ca. 1500 m , Dwyer 7037 (MO), Dwyer & Hayden 7686 (MO), Lewis et al. 642 (GH, MO), Pittier 2943 (GH, NY, US); Bajo Mono, Davidson 469 (A, F, MO, US); Cerro Horqueta, Blum & Dwyer 2608 (MO); Finca Lerida to Boquete, Woodson et al. 1106 (A, MO, NY); vicinity of Puerto Armuelles, Woodson & Schery 828 (MO); vicinity of Progreso, Cooper & Slater 289 (US), Cooper & Slater 187 (F, GH, NY, US, Y). Code: Cocoli, Riley 133 (MO, US); between Las Margaritas & El Valle, Woodson et al. 1726 (A, F, MO, NY). Herrera: road between Las Minas and Prese, Duke 12335 (MO); 12.5 miles S of Ocu, Lewis et al. 1668 (GH, MO, UC, US); Ocu, Ebinger 2002 (MO). Los Santos: Guayabo, W of Tonosi, Stern et al. 1903 (MO): Pediasi, Dwyer 2483 (MO); 0.5 miles N of Tonosi, Tyson et al. 3108 (MO). Panama: thickets and forest near Arraijan, Woodson et al. 1367 (A, F, M O , N Y ) ; Chepiho Island, Duke 10317 (MO); ca. 5 miles N of Panama City, Correa et al. 5045 (MO); Pedro Gonzalez, Perlas Island, Allen 2594 (MO); 3 miles E of Panama City, Maxon et al. 7083 (NY, US); Taboga Island, Standley 27946 (US), Woodson et al. 1436 (A, F, M O , N Y ) ; Tocumen, Dwyer 1850 (MO), 4065 (MO). Veraguas: Chiriqui-Ver aguas boundary on old gravel road, Dwyer 2930 (MO); ca. 5 miles N of Santiago, Blum & Tyson 607 (MO); Santiago, 4 miles toward Atalaya, Dwyer & Kirkbride 7419 (GH, M O , US); roadside savanna 2-4 miles E of Santiago, Duke 12357 (MO). COLOMBIA. Antioquia: Fredonia, 1850 m. Archer 517 (US); vicinity of Dabeiba, ca. 1350 m , Barkley & Gutierrez 1783 (NY, US); vicinity of MedeUin, Archer 772 (US), Delisle & Gutierrez 30 (F, US), Torro 444 (US); bank of Rib Magdalena, Pennell 3706 (GH, N Y ) , Pring 16 (MO). Caqueta: near Florencia, Little 9123 (US). Choco: Andagoya, 70-100 m , Killip 35072 (US). Cundinamarca: Caparrapi, ca. 1275 m, Barriga 7640 (US); El Colegio, Joseph 1047 (US), Dugand & Jaramillo 2950 (US); vicinity of Bogota, Vogel 78 (VEN); E of Guaduas, Barriga 11731 (US); 11 k m E of Gutierrez, Grant 9785 (US); La Mesa, 1350-1400 m , Fernandez & Mora 1305 (NY); between El Salto & Las Mesitas, Dugand & Gutierrez 3520 (US); La Vega, Barriga 10642 (US), Arbelaez & Cuatrecasas 5343 (F). Huila: Finca Cedral, Rib Fortalecihas, Fosberg 19730 (US), 19736 (US); vahey or Rib Neiva, between Algeciras and Salado, Fosberg 19303 (GH, US); road between Garzon and Florencia, Mason 13901 (US); Pitalito, open fields, Schultes & Villarreal 5104 (F, GH, MO). Meta: Uribe, Rib Duda, Fosberg 19465 (US); Villavicencio, Cuatrecasas 4548 (F, US). Putumayo: Mocoa, 750-850 m, Schultes & Smith 3000 (GH). San Andres: San Andres Island, Proctor 3316{US). Santander: Bucaramangaand vicinity, Killip & Smith 16191 (A, G H , N Y , US); between Bucaramanga and El Jaboncillo, Killip & Smith 16364 (US). Norte de Santander: vicinity of Chinacota, Killip & Smith 20784 (A, US); between Chinacota and La Esmeralda, Killip & Smith 20915 (A, US); El Ceibal, Rib Margua, Cuatrecasas 13422 (US). Valle: CuchiUa, E of Zarzal, Pennell et al. 8534 (GH); Canta Claro, Figueiras 8071 (US); Rib Cauca, Sneidern 4560 (F); Quebrada Robada, Rib Alban, Cuatrecasas 22394 (F); Cauca valley, Zarzal, Killip 8353 (GH, NY). VENEZUELA. Apure: San Fernando de Apure, Trujillo 2195 (VEN). Aragua: road to Choroni, Ramia 716 (VEN); vicinity of Maracay, Burkart 16276 (VEN), Doreste 51 (VEN), Standen 80 (VEN), 96 (VEN); near Ocumare and the coast, Badillo 1830 (VEN); La Victoria, H o t 285 (VEN). Carabobo: Tocujito, Saer 895 (NY, US). Federal District: Cerros del Avila, 1600 m, Pittier 39 (NY); upper Cotiza near Caracas, Pittier 7193 (GH, US); between Caracas and La Guaira, Rose & Rose 21697 (US); near Guaira, Badillo 3009 (VEN); above Caracas, between Los Vanados and Papelon, Steyermark 55063 (A, F, US); Coticita, Curran & Haman 1058 (GH, N Y , US); Biscaina, Fendler 591 (GH, N Y ) ; San Bernardino, Fernandez 189 (VEN); Rib Vahe, El Valle, Fernandez 182 (VEN). Merida: Mese Bolivar, Bernardi 638 (NY); along road to La Carbonera, 36 k m W of Merida, Breteler 3133 (VEN); Ejido, Little 15879 (VEN); Puente La Victoria, Bernardi 513 (NY). Miranda: Chacao, 960 m, Williams 12164 (A, F); Las Mostazas beyond Teques, Allart 192 (NY); Los Teques, 1400-1500 m, Pittier 7517 (GH, US). Portuguesa: road to Acarigua, Chardon 105 (VEN); Guanare, Trujillo 1825 (VEN); road to Piritu, Trujillo 1765 (VEN). E C U A D O R . Bolivar: valley of Rib Playas, 3-5 km above Balzapamba, Fosberg 22507 (US); vicinity of Pucara, Solis 6991 (F); vicinity of Tehmbela, Solis 6886 (F). Chimborazo: vicinity of Bucay, Solis 5175 (F); S bank of Rib Chimbo, N of Sacramento region, Wiggins 11109 (F, US); between Huigra and Naranjapata, Hitchcock 20685 (GH). Cotopaxi: bank of Rib Macuchi near Macuchi, Steere 8089 (NY). Esmeraldas: Lita, ca. 500 m, Jdtiva & Epling 858 (NY). Guayas: along stream near Pedro Carbo, Haught 3028 (F). Loja: Rib Chiguango, 13 k m WNW of Catamayo, Fosberg & Giler 22961 (NY, US). Napo Pastaza, vicinity of Puyo, Skutch 4516 (A, F, US). El Oro, between Rib Minas Nuevas NW to above Paccha, Steyermark 54128 (F). Province unknown: vicinity of Ventura, Rose & Rose 23516 (GH, US). PERU. Amazonas: Chachapoyas, Mathews 3135 (GH, NY); vicinity of Campomiento Ingenio, 1300-1400 m , Hutchison & Wright 3826 (MO, UC, US); rain forest along Rib Marahon, 2-10 k m above mouth of Rib Santiago, Wurdack2258 (NY, US); along Rib Utcubamba, ca. 40 k m S of Bagua Grande, Hutchison 1464 (NY, UC); Rib Utcubamba, between Bagua Grande and Campomiento Ingenio, Hutchison 4452 (MO, UC, US). Cajamarca: Santa Rosa, 1200 m, Woytkowski 5709 (MO, GH). Cuzco: QuiUabamba, 1040 m. Stork et al. 10447 (A, F); Potrero, 1300 m, Nasgarto 1161 (GH). Huanuco: between Huanuco and Pampayacu, Kanehira 197 (A); vicinity of Tingo Maria, Dwyer 6269 (MO), Ferreyra 1587 (US), Stork & Horton 9513 (A), vahey of Rib HuaUaga, Belshaw 3076 (US), Rio Tulumayu effluent of Rio HuaUaga, Fosberg 28835 (F); Pozuzo, ca. 650 m, MacBride 4679 (F, GH, US). Junin: Colonia Perene, dense forest, Killip & Smith 24966 (GH, US); Chanchamayo, Soukup 2906 (F); between La Merced & Quimiri, Ferreyra 3695 (US); La Merced, MacBride 5416 (F, GH, US) 5417 (GH, US), Killip & Smith 23814 (F); Rib Negro to Satipo, Woytkowski 5804 (GH, MO, US); without exact locality, Evinger 538 (NY). Loreto: Balsapuerto, ca. 220 m, Klug 2928 (A, F, GH, MO, NY, US); lower Rio Huallaga, Williams 4785 (F); 15 k m N of Iquitos, Dodson 2858 (MO); Mishuyacu, near Iquitos, Klug 521 (F, US); Pevas, at mouth of Rib Ampiyacu on Rib Amazonas, Fosberg 29222 (F); Pucalipa, 200 m, Woytkowski 6276 (GH, M O , US); Rib Putumayo, at mouth of Rib Zubineta, forest, Klug 2062 (A, F, GH, MO, NY, US), 2219 (F, GH, NY, US); near Yurimaguas, Spruce 3890 (NY). Piura: Canchaque, 1150 m. Stork 11399 (GH). San Martin: HuinguiUo, forest, 500 m, Woytkowski 7137 (GH, MO); Lamas, 840 m, Williams 6336 (F); near Moyobamba, forest, Klug 3302 (A, F, GH, MO, NY, US); Soritor, forest 900 m, Woytkowski 6242 (GH, US); in jungle on ridge E of Tingo Maria, Allard 22414 (F, NY); vicinity of Tarapoto, Ferreyra 7754 (US), Spruce 4188 (GH, US), Williams 5499 (F), Woytkowski 35019 (MO). Province unknown: without exact locality, Mathews 3135 (GH), s.n. (NY), Spruce s.n. (NY). BRAZIL. Federal District: ca. 25 k m N of BrasUia, 950 m, Irwin et al. 11295 (MO, NY). Minas Geraes: Caldas, Regnell 1043 (US); Ituiutaba, Macedo 4511 (US); Vicosa Barbado watershed, 690 m, Mexia 4591 (A, F, GH, MO, NY, US); without exact locahty, Pires & Black 3066 (NY). Parana: at Iguazu, banks of Rib Parana, Beetle 2131 (US); National Park of Iguazu, Pereira 5319 (F); Therezina, Dusen s.n. (F, GH). Rib Branco: Acre territory, Wilde & Baldwin 18 (A, F). Sao Paulo: Sao Simao, Hoehne 11533 (F); Ytu, Hoehne 31422 (GH, NY, US). Santa Catarina: Itapiranga, ca. 200 m. Smith & Klein 11809 (NY). BOLIVIA. Cochabamba: Riberaha, 400 m, Cardenas 4165 (US); vicinity of Cochabamba, Bang 1222 (A, GH, MO, NY, US). La Paz: near Coroico, Polo-Polo, Buchtien 5768 (GH, US); Huachi, head of Rib Beni, White 457 (F, NY); vicinity of Maripi, Buchtien 1626 (US), Rusby 2105 (F, NY), Williams 803 (NY); MiUuguaya (?), Buchtien 4004 (GH, NY). Santa Cruz: Angostura, Cardenas 4642 (US), Steinbach 294 (F); vicinity of Santa Cruz, Cardenas 5774 (US); Santa Cruz de la Sierra, Guarayos mission, Werdermann 2664 (MO); La Cruz, Kuntze s.n. (US). ARGENTINA. Misiones: Catarantas del Iguazu, Descole 3204 (NY). Saka: Boca de la Quebrada, 500 m , Schreiter 3543 (A); Embarcacion, 450 m, Schreiter 203 (F), Venturi 5156 (A, F, US); Iquira a Aguaray, 560 m , Schreiter 11177 (F, GH); Rib Pescado, Campamento, Meyer 4813 (F); Tranquita-Tartagel, Schulz & Varela 5160 (A); Vespucio, Maldonado 807 (F). PARAGUAY. Amambay: Sierra de Amambay, Rojas 10283 (NY). Concepcion: Rib Aquidaban, Fiebrig 4260 (GH). Alto Parana: without exact locality, Fiebrig 5661 (US), Bertoni 1343 (A). San Pedro: Puerto Antequera, forest trail, Rojas 2342 (F). Province unknown: without exact locality, Hassler 5212 (NY), 6854 (A, MO, NY), Jorgensen 4313 (A, GH, NY).

Discussion:Hamelia erecta Jacq., Enum. Fl. Carib. 16. 1760 (holotype, Jacquin s.n.). H. coccinea Swartz, Prodr. 46. 1788 (holotype, Swartz s.n.). H. sphaerocarpa Ruiz & Pav., Fl. Per. & Chil. 2: 69. t. 221. 1799 (holotype, Ruiz & Pavon s.n. MA?). Duhamelia patens (Jacq.) Pers., Synop. PL 1: 203. 1805. D. sphaerocarpa (Ruiz & Pav.) Pers., Synop. PI. 1: 203. 1805. Hamelia patens y ?quinifolia D C , Prodr. 4: 442. 1830 (holotype, Mocino & Sesse s.n. MA?). H latifolia Reichb. ex D C , Prodr. 4: 442. 1830 (type, Sieber 32). H. lanuginosa Mart. & Gal., Bull. Acad. Brux. 11: 233. 1844 (holotype, Galeotti 2615, P, F frag.!). H. corymbosa Sesse & Mociiio, La Nat., ser. 2, 1, app, 37. 1888 as Hamellia (type, Mocino & Sesse s.n., MA?). H. brittoniana Wernh., Jour. Bot. London 49: 346. 1911 (holotype, Kuntze 2178, NY!). H. intermedia Urban & Ekman, Arkiv Bot. 24A(4): 46. 1932 (holotype, Ekman 10857, S; isotype A!). The very common Hamelia patens var. patens is easily recognized by villous pubescence, and orange-red tubular corolla, ovate calyx lobes, and sessile to shortly pedicellate flowers secund on a helicoid axis. This variety is closely related to H versicolor and H. sanguinea, both of which may have been derived from H. patens. The degree of pubescence in var. patens varies from sparsely villous to densely villous. This is especially evident when populations from southern Florida, the West Indies, and northern South America are compared with populations from Mexico. Most Mexican specimens examined had a very dense indument over most of the plant. As a result, the more pubescent populations in Mexico had been described as H. lanuginosa. A complete gradation from the densely to the sparsely villous condition is found in the Central American specimens. The specimens examined from the remainder of the range of var. patens had only sparse, long, soft and often curled trichomes on the leaves, stems and flowers. Hamelia patens and H. erecta were described at the same time by Jacquin (1760); later investigations have shown that both names refer to the same species. Since H. erecta preceded H. patens by three lines in the initial publication, it might be concluded that H. erecta is the legitimate name. This name was accepted by Standley, and H. patens was placed in synonymy with H. erecta in his Trees and Shrubs of Mexico. A few years later in the North American Flora he reversed the two names, holding that H. patens was the correct name. According to Article 57 of the International Code of Botanical Nomenclature, the choice of the person w h o first unites taxa bearing names or epithets of the same date must be followed. A thorough search of the literature was undertaken to find this author. As far as I have been able to determine, L'Heritier (Sert. Angl. 4. t. 7. 1788), who considered H. erecta to be a synonym of H. patens, was the first to publish this, although the problem may never be resolved with absolute certainty.
Distribution:United States of America North America| Bahamas South America| Jamaica South America| Haiti South America| Dominican Republic South America| Puerto Rico South America| Guadeloupe South America| Virgin Islands South America| Dominica South America| Martinique South America| Mexico North America|