Monographs Details: Mielichhoferia
Authority: Sharp, Aaron J., et al. 1994. The Moss Flora of Mexico. Part One: Sphagnales to Bryales. Mem. New York Bot. Gard. 69 (1)
Family:Bryaceae
Scientific Name:Mielichhoferia
Description:Genus Description - Small, slender plants in dense, yellowish or reddish, ± shiny tufts consisting mainly of subjulaceous subfloral innovations. Leaves of innovations erect, crowded, concave, ovate to ovate-lanceolate or oblong-lanceolate, unbordered; margins entire to indistinctly serrulate; costa strong, percurrent or nearly so, rarely short-excurrent; cells lax and thin-walled, long-rhomboidal, becoming short-rectangular below. Dioicous. Perichaetia basal, overtopped by subfloral innovations and thus appearing lateral Setae slender, flexuose, curved at the tip; capsules mostly pendent, symmetric, mostly inflated-pyriform, with a small and a ± short, broad neck; annulus broad, revoluble; operculum conic or convex-conic, blunt; stomata superficial; peristome single and consisting of exostome only, or double with a much reduced endostome, the exostomial teeth evenly spaced or approximate in pairs, lanceolate, variously roughened.

Discussion:

In this treatment, Haplodontium has been included in Mielichhoferia, and the species traditionally assigned to Mielichhoferia are given the name Schizymenium. Haplodontium has been defined in terms of single exostomial peristomes contrasting with single endostomial peristomes of Mielichhoferia. Yet, the peristomes that have been allowed within Mielichhoferia vary from single and endostomial or exostomial to double. The peristome ofthe type species ofMielichhoferia, M . mielichhoferiana (Funck ex Hook.) Loeske, though seemingly single and exostomial, actually consists of a a more or less well-developed exostome, as well as a delicate, somewhat adherent endostomial basal membrane, not unlike the peristomes that have been allowed within Haplodontium. Thus, it is impossible to define either genus on the basis of peristome structure. Mielichhoferia, in this sense, can be characterized by dioicous inflorescences and peristomes that are single and exostomial or double with a much reduced endostome, with exostome teeth generally roughened. The name Schizymenium can be apphed to monoicous species, with peristomes single and endostomial, or rarely double or lacking, with exostome teeth, if present, smooth.

For further information on the relationships and nomenclature of the subfamily Mielichhoferioideae, see Crum and Shaw (1982) and Shaw (1984,1985).