Monographs Details: Montanoa mollissima Brongn.
Authority: Funk, Victoria A. 1982. The systematics of Montanoa (Asteraceae, Heliantheae). Mem. New York Bot. Gard. 36: 1-133.
Description:Species Description - Shrubs 1-2(-4) m tall. Stems terete, brown, herbaceous parts puberulent. Leaves variable; petioles 1-6 cm long, partially or completely winged with auricles at base, puberulent to moderately pubescent, hairs 0.5-1.0 mm long; blades lanceolate to triangular (Fig. 36A-G), 2.5-14.5 cm long, 1-10 cm wide, apex acute to acuminate, margin entire to irregularly dentate to 3-lobed, adaxial surface densely pubescent, hairs pustular, abaxial surface densely glandular and pubescent, hairs white, less than 1.0 mm long. Peduncles 1.5 cm long, densely puberulent. Heads erect, 1.0-1.2 cm diam in flower, 2.0-2.5 cm diam in fruit, several to many in open cymose synflorescences in oppositely branched compound corymbs. Phyllaries 6-8, more or less uniseriate, equal, ovate-lanceolate, 4-7 mm long, 1.5-2.0 mm wide, green, apex acute to acuminate, margin ciliate, entire, abaxial surface densely pubescent, hairs 1.0 mm long, adaxial surface essentially glabrous. Ray florets 8-12; corollas white, ligules 15-30 mm long, 6-8 mm wide, adaxial surface essentially glabrous, abaxial surface moderately glandular and sparsely pubescent especially on the veins, hairs less than 0.5 mm long, tube 1.0-1.5 mm long, 0.5 mm wide, glabrous. Disc florets (Fig. 36H) 75-100; corollas yellow, tube 1.0-1.5, 0.5 mm diam, glabrous, throat cylindrical, 2.0 mm long, 1.0 mm diam, essentially glabrous, lobes 5 (rarely 4), 1.0 mm long, 0.75 mm wide, apex acute, densely glandular and pubescent, hairs thick, less than 0.5 mm long; stamens with filaments 2.0 mm long, 0.2 mm wide, anthers not fully exserted from corolla, thecae yellow to brown, 1.5-1.75 mm long, 0.5 mm wide, apical appendages yellow, acute, 0.5 mm long, 0.5 mm wide, abaxial surface of apical appendage glandular, styles yellow, 4.5-5.5 mm long, enlarged at base, stigmatic surface yellow, 1.0 mm long, apical appendage yellow, long-acuminate, 0.5 mm long, 0.25 mm wide. Pales at anthesis ovate-triangular (Fig. 36I) 4-5 mm long, 1.0 mm wide, light yellow, apex long-acuminate, indurate, yellow, abaxial surface sparsely glandular and pubescent, hairs 1.0 mm long, adaxial surface glabrous, margin ciliate, entire; pales at fruiting persistent, almost narrowly triangular (Figs. 32A; 36J, K), 9-12 mm long, 2.5-3.0 mm wide, indurate with prominent parallel veins, stramineous, apex tapering to a slightly recurved and indurate tip, margin sparsely ciliate, entire, abaxial surface sparsely glandular and pubescent, hairs 0.5-1.0 mm long, adaxial surface glabrous. Achenes brown-black, 3.0-3.5 mm long, 2.0 mm wide, smooth. Chromosome number, n = 19.
Flowering (and Fruiting) Period: July-September (August-November).
Discussion:Type: MEXICO, collected from Hort. Paris, 1851 from plant “grown from seeds sent from Mexico by A. B. Giesbreght in 1843,” A. B. Giesbreght s.n. (holotype, P!; isotypes, FI! P!; photos of holotype, GH! OS! US!; fragment of holotype, GH!). Eriocoma mollissima (Brongniart ex Groenland) Kuntze, Rev. Gen. Pl. 1: 336. 1891.
Montanoa pringlei Robins. & Greenm., Proc. Amer. Acad. Arts 34: 512. 1899. Type: MEXICO, Puebla, calcareous hills near Tehuacán, 28 Jul 1897, C. G. Pringle 7491 (holotype, GH!).
Montanoa tehuacana Robins., Proc. Amer. Acad. Arts 47: 209. 1911. Type: MEXICO, Puebla, tufa bluffs near Tehuacán, 1680 m, 7 Aug 1901, C. G. Pringle 8585 (holotype, GH!; isotypes, BM! C! CM! CU! F! FI! G! GOET! K! LE! M! MEXU! MICH! MO! MSC! NY! P! S! UC! US! W!; photo of isotypes from K and P, OS!).
A much-branched shrub that usually appears straggly because of the relatively small leaves (blades not over 14.5 cm long), that are often few and widely spaced on the branches. The species is distinct because of its habit and completely or nearly completely winged petioles. Montanoa mollis sima is most closely related to the sub-cloud forest tree, M. guatemalensis with which it shares a white abaxial leaf surface, 70-100 disc florets and similar mature pales. But, it is easily distinguished from this species by the presence of winged petioles and the shrubby habit. Also, the mature pales of the two species, although similar in shape and size, differ in that those of M. guatemalensis are more recurved (cf. Figs. 36J, K; 35L).
Plants of this species (e.g., Cronquist & Sousa 10394) have been collected growing with members of M. tomentosa subsp. xanthiifolia (e.g., Cronquist & Sousa 10395) near Tehuacán where both taxa are common. No morphological evidence for hybridization has been found; the two species appear distinct in all collections from that area.
Distribution:Mexico North America