Tocoyena guianensis var. communis Steyerm.
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Authority
Maguire, Bassett. 1965. The Botany of the Guayana Highland. Mem. New York Bot. Gard. 12 (3): 1-285.
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Family
Rubiaceae
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Scientific Name
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Type
Type. Small tree 5 m tall, 6 cm diam; corolla white; shade at margin of river, Rio Oiapoque, opposite Pedro Alice, 3° 40' N, 52° 1' W , Territorio Amapa, Brazil, 18 Aug 1960, H. S. Irwin, J. M. Pires & L. Y. Th. Westra 47620 (holotype NY). Paratypes. BRAZIL. Amapa: Rio Araguari, along river between Camps 5 and 6 and tin mine trail, 1° 26' N, 51° 58' W , and 1° 9' N, 51° 52' W , Pires, Rodrigues & Irvine 50788; Rio Araguari, varzea forest. Camp 13, 1°'45' N, 25° W, Pires, Rodrigues d:. Irvine 51600. FRENCH GUIANA. Rio Oiapoque, between Pedro Alice, 3° 40' N, 52° 1' W , French Guiana, and Roche Mon Pere, 3° 33' N, 52° 5' W, Brazil, Eejler 47640. Crique Alikene, about 50 km from its confluence with River Camopi, ca. 3° 10-20' N, 52° 28-32' W , Pires 48580; about 1 km west of Cachoeira Utussansain, 2° 8' N, 52° 55' W, Irwin, Pires d Westra 48072. VENEZUELA. Territorio Federal Amazonas: Rio Siapa just above Raudal Gallineta, Wurdack & Adderley 43532.
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Discussion
A specimen from French Guiana (Irwin, Pires & Westra 47993 from Cachoeira Utussansain) is a more scabridulous-leaved form, in which the upper surface of the leaf-blades are more scabridulous, and the nerves of the upper leaf surface are more densely strigose; the petioles also are more densely hirtellous-scabridulous.