Authority: Berg, Cornelius C. 2001. Moreae, Artocarpeae, and (Moraceae): With introductions to the family and and with additions and corrections to Flora Neotropica Monograph 7. Fl. Neotrop. Monogr. 83: 1-346. (Published by NYBG Press)
Discussion:In general the subspecies recognized in the monograph can still can be told apart, even in the northwestern part of the Amazon Basin where all subspecies, except for subsp. castilloides, co-occur (sometimes even in the same locality). In a small number of collections features are mixed, possibly as a result of hybridization.