Monographs Details: Fissidens sublimbatus Grout
Description:Species Description - Plants usually olive green, rusty-red in stems and costa, often partially covered with soil. Stems often dimorphic, usually branched from older parts, to 6 mm long x l.5 mm wide, infertile ones longest; rhizoids basal and axillary, smooth, tan to brown-red; axillary hyaline nodules absent; epidermis and outer 1-2 tiers of cortical cells small, incrassate, usually pigmented; inner cortical cells larger, thin-walled, hyaline; central strand absent. Leaves inrolled when dry, imbricate to distant, as many as 22 pairs on infertile stems, usually fewer on fertile stems, oblong to lanceolate, rounded to broadly acute to obtuse-apiculate, apiculus formed of a single, clear, sharp cell, to 1.0 mm long x 0.10.4 mm wide, perichaetial and larger leaves oblong to oblong-ligulate and lanceolate, to 1.3 mm long x 0.20.3 mm wide; margin ± entire, serrulate distally, elimbate or limbate, limbidial cells unistratose, restricted to vaginant laminae or found to variable degrees on all laminae, often conspicuously intralaminal in proximal parts of vaginant laminae; costa ending 2-5 cells below apex or percurrent in perichaetial and larger leaves, bryoides type; dorsal lamina usually narrowed abruptly, often arched, ending well above insertion; vaginant laminae of cauline leaves 2/3-3/4 leaf length, acute, ± equal, 1/2-3/4 leaf length in perichaetial leaves; laminal cells ± obscure, eguttulate, unistratose, smooth, bulging, subquadrate, 6-10 µm long, in transverse section twice as deep as wide, often arranged in discernable rows in dorsal and ventral laminae. Monoicous (cladautoicous, gonioautoicous, possibly rhizautoicous); perigonia and perichaetia terminal on stems and branches or perigonia gemmiform, axillary. Sporophytes 1 per perichaetium, reddish; seta smooth, to 5 mm long; theca exserted, 0.5-1.0 mm long, slightly inclined, bilaterally symmetric to erect, radially symmetric, stomatose, exothecial cells quadrate to oblong, vertical walls thicker than horizontal walls; peristome bryoides type; operculum conic, short-rostrate, to 0.3 mm long. Spores smooth, 14-23 µm diam. Calyptra cucullate, smooth, to 0.6 mm long.
Discussion:Fissidens sublimbatus is closely allied to F. crispus. Both species have similar laminal cells, but F. sublimbatus differs in its restricted limbidium, a dorsal lamina that in most leaves ends well above the leaf insertion, the lack of axillary hyaline nodules, a shorter costa, and a leaf apex that in many leaves ends in a single, clear, sharp cell.
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