Monographs Details: Tococa bolivarensis Gleason
Authority: Michelangeli, Fabián A. 2005. (Melastomataceae). Fl. Neotrop. Monogr. 98: 1-114. (Published by NYBG Press)
Description:Species Description - Subprostrate to procumbent shrub, to 1.0-1.5 m tall, the branches up to 2.5 m long, with adventitious roots developing on the portions closer to the ground; stems with a mix of short glandular setae 0.2 mm long and stellate hairs, caducous, the nodes glabrous. Leaves isophyllous, elliptic to ovate, 2-5 X 1.2-3.0 cm, apex broadly acute to obtuse, base rounded to broadly acute, adaxial surface sparsely glandular setose, sometimes the trichomes disposed in lines parallel to the secondary veins, abaxial surface sparsely glandular-pubescent, densely hirsute on the primary and secondary veins, light green, 3(-5)-nervate, the veins impressed on the adaxial surface, the tertiary veins not evident, coriaceous, entire or ciliolate, the setae up to 1.8 mm long, sometimes revolute; petioles 0.3-1.0 cm long, glandular-setose; domatia absent. Inflorescence a terminal cyme of 2-4 flowers, 1-3 cm long, the axis dark red to magenta, terete. Flowers on pedicels 1-3 mm long, flared at the apex, subtended by bracts persistent until anthesis, elliptic, 0.7-1.3 X 0.3 mm, glandular-setulose; hypanthium globose, with the base as wide as the pedicel, 4.4-5.0 mm long, sparsely glandular-setulose; outer calyx teeth reduced and inconspicuous, sometimes with a short apical glandular seta or a sessile gland, 0.2 mm long, inner calyx teeth deltoid, fused at the base, 1.0-1.5 mm long; the ring inside the torus glabrous; petals 5, ob-ovate, 6.5-11 X 5.0-6.5 mm, base attenuate, apex emarginate, slightly pruinose, glabrous, red to dark pink; stamens all the same size; filaments 5-6 mm long, glabrous; anthers yellow; the connective with a dorsal-basal blunt tooth; thecae 5 mm long, opening by a slightly ventrally inclined pore; ovary 3-locular, 1/3 inferior, the superior portion terete, the apex truncate; style glabrous, 12-14 mm long; stigma truncate. Fruits globose, 4.5-6 mm long, black, glabrous, with the calyx persistent and slightly reflexed; seeds not evident through the pericarp, truncate-obovate, 0.40.5 mm long, without capitate trichomes in the raphal area, without sculpturing, anticlinal walls straight, periclinal walls flat, boundaries between periclinal walls flat.

Discussion:Tococa bolivarensis is a creeping shrub endemic to the eastern Guayana Highlands. Although it has been collected only in Venezuela, it probably occurs also on the Brazilian side of Serra do Sol and in Guyana on the slopes or summit of Roraima. Even though there are only a handful of collections, it is locally abundant in the summit of Auyán-tepui and can be easily distingished from other tepui Melastomataceae by its ciliolate leaves and reddish thick petals. For alliances and similarities, see discussion under T. raggiana.
Distribution:Bolívar Venezuela South America|