Monographs Details: Comarostaphylis spinulosa subsp. glandulifera Diggs
Authority: Luteyn, James L., et al. 1995. Ericaceae, Part II. The Superior-Ovaried Genera (Monotropoideae, Pyroloideae, Rhododendroideae, and Vaccinioideae P.P.). Fl. Neotrop. Monogr. 66: 560. (Published by NYBG Press)
Description:Subspecies Description - Low, often densely branched shrubs to 2 m tall. Bark furrowed and peeling, gray to orange-gray; young twigs sometimes red, glandular hirtellose to hirsute with stalked, swollen-headed glandular trichomes to 2 mm long, with short, uniseriate, hyaline hairs also present. Leaves coriaceous, elliptic to obovate or ovate, 2-7.4 × 0.8-3.7 cm, marginally serrate to serrulate, upper surface glabrate or with scattered glandular or eglandular trichomes, lower surface with dense white to ferruginous tomentum imparting a distinct color to the surface, also with multiseriate-stalked, swollen-headed glandular trichomes, especially along the midrib; petioles with indumentum as on young twigs. Inflorescences terminal, racemose or paniculate, usually with few branches, 3.3-9.2 cm long, rachis, pedicels, and bracteoles densely glandular hirtellous to hirsute with stalked, swollen-headed glandular trichomes to 1.5 mm long, shorter hyaline hairs also present; floral bracts lanceolate to narrowly ovate, flattened to slightly navicular, 3-ll(-20) × 0.4-2.5(-6) mm, tomentulose by eglandular trichomes, also with glandular trichomes, glandular-ciliate; pedicels 0.4-1 (-1.4) cm long, bracteoles basal to medial, lanceolate, 2.5-5 mm long, glandular-ciliate. Flower buds sometimes red; calyx lobes triangular to broadly so, 1.5-2.1 × 1.3-1.4 mm, acute, glabrous to densely pubescent with glandular or eglandular hairs, the trichomes often more dense at base of lobes, ciliate or not.

Discussion:The abundant glandular trichomes and the much denser tomentum on the abaxial leaf surfaces serve to distinguish subsp. glandulifera from subsp. spinulosa. Because of the glandular pubescence, specimens of this subspecies have in the past been confused with the quite different C. longifolia of central to W Mexico despite the larger overall size (to 8 or possibly even 15 m tall), much larger (>8 cm long) lanceolate to elliptic leaves, and longer, more branched inflorescences of the latter.

Distribution and Ecology: (Fig. 17) and ecology. Subspecies glandulifera is known only from N Oaxaca and extreme S Puebla, with the exception of one collection (MacDougall s.n., NY) from south-central Oaxaca. The typical habitat is moist Quercus-Pinus-Arbutus forests with Quercus brachystachya, Q. obtusata, Pinus pseudostrobus, Arctostaphylos pungens, Gaultheria sp., Pernettya prostrata and Vaccinium sp. at 2000-2500 m elevation. Flowering Jun-Feb; fruiting Jun-Feb.

Distribution:Mexico North America| Oaxaca Mexico North America| Puebla Mexico North America|